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Revealed: How the fish got its fingers By Steve Connor, Science Editor 
02 April 2004


A two-lane highway in America has helped scientists to explain one of
life's most enduring mysteries: how fish grew the fingers that enabled
them to crawl out on to land.

The road in Pennsylvania happened to be cut out of 365 million-year-old
rock in which the researchers found the oldest known fossilised arm bone
of one of the world's first four-legged creatures, or tetrapods.

Specialists said that the bone - a prototype humerus - can help to
explain how ancient fish with primitive lungs grew rudimentary limbs from
their fins which allowed them to make the first tentative steps towards a
terrestrial life.

The discovery, in the journal Science , suggests the first tetrapod limbs
were used to prop up the heads of air-breathing fish and only afterwards
did the fish begin to use these legs to clamber out of water. The
fossilised bone is thick and flat and would not have allowed much
movement between the limb and the shoulder, indicating that it was of
little use other than to prop up the creature's front half.

Michael Coates, of the University of Chicago, said: "When this humerus is
compared to those of closely related fish, it becomes clear that the
ability to prop the body is more ancient than we previously thought. This
bone is a lot more robust than a humerus from any of the ancient species.
Relative to other tetrapods, this is almost over-engineered. There's a
massive space for the attachment of substantial muscle going to the
chest.

"This means that many of the features we thought evolved to enable life
on land originally evolved in fish living in aquatic ecosystems."

The tetrapod creature probably lived in shallow, muddy water during the
late, or Upper Devonian period between 360 million and 370 million years
ago when the exploration of dry land by vertebrates had yet to start. At
that time, the land was largely the domain of insects, rather than
animals with backbones, and the water was occupied by fish with bony
skeletons and fins with hand-like bone structures that were to evolve
into vertebrate limbs.

Dr Coates and his colleagues Neil Shubin and Edward Daeschler believe the
fossilised bone found in Pennsylvania helped the forelimb fulfil an
intermediate function between the braking and steering of a fish's fin
and the walking movements of an early amphibian.

Drs Daeschler and Shubin found the fossil in 1993 when they were
excavating near the highway but it took nearly eight years to discover it
was important.

"We found a number of interesting fossils at the site," Dr Daeschler
said. "But the significance of this specimen went unnoticed for several
years because only a small portion of the bone was exposed and most of it
lay encased in a brick of red sandstone."

After the bone was fully excavated, its true significance quickly became
apparent, Dr Shubin said. "We knew it was a humerus, but it was an
entirely different kind. We had never seen one like it before. It's a
mosaic of primitive fish and derived amphibian."

Jenny Clack, a vertebrate palaeontologist at Cambridge University, said
the primitive forelimb could have propped the creature's head out of
water to allow it to breathe air, or it could have been used to anchor
the animal in fast-moving water, Dr Clack said. "It begins to fill in the
picture about what we think about the transition. The difference between
fish and these early tetrapods is becoming increasingly blurred."

The same palaeontology site in Pennsylvania has yielded two other types
of tetrapod living in the Devonian period, Dr Clack said. "If this is
really a third form, it hints at a wide diversity of tetrapods existing
in close proximity, in what is emerging as one of the richest and most
varied of any late Devonian vertebrate site," she added.

The scientists who have excavated the Pennsylvania site said it contains
fossils of other plants and animals that suggest the area was "teeming"
with life more than 360 million years ago. 

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