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Andreas,

Is Phaser putting your dimer search model directly on the crystallographic
2-fold?  This will generate a very large number of clashes between monomer
1 and monomer 2 during the packing routine.  The simplest way around this
is to re-run MR_PAK after trimming your search model to 1 monomer.  If the
dimer search model is not placed directly on the crystallographic 2-fold,
does your data demonstrate a large patterson peak (other than the
C-centering)?

Eric

On Thu, 16 Nov 2006 [EMAIL PROTECTED] wrote:

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> 
> 
> Dear molecular replacement cracks out there,
> 
> 
> I'm working on a non-trivial molecular replacement problem (bad model,
> poor data, many molecules in au) and am hitting the wall.  This might be
> because the problem cannot be solved or because I'm blind to the solution.
> 
> Run on a list of normal mode-perturbed models, Molrep didn't find obvious
> solutions but suggested a dimer I should use as model instead of the
> monomer.  This makes good sense, as my protein is like dimeric (though I
> can't see NCS in the data).  What's the basis of this suggestion, Alexei?
> 
> Using the dimer didn't help Molrep find the solution, but Phaser suggested
> a rotation function (Z-score = 3) whose 26 associated translation
> functions  have Z-scores between 7 and 9.  All solutions are rejected
> because of clashes (~100).
> 
> Forcing the transformations I can write out 26 potential solutions and
> look at them in PyMOL.  They are lined up along the a axis of the unit
> cell (c222_1). The a axis is also coincident with the two-fold that
> relates the two monomers of the dimer (thus the nearly complete overlap
> indicated by 100 clashes).  One monomer is symmetry-related to the other.
> 
> Why does Phaser put the dimer on a symmetry axis?  Would this not give
> wrong solutions in any case where you search with a dimer?  Is it possible
> (and sensible) to define the a axis in C222_1 as a dyad axis for molecular
> replacement?
> 
> At first I thought my problems were due to misindexing, but running phaser
> or Molrep with data reindexed in all other possible spacegroups (p1, p2,
> p2_1, c2, c222, see statistics at the end of the email) didn't give
> solutions (nor high Z-scores in Phaser).  For example, in P1, Molrep
> places 4 dimers but the contrast values decrease with every placed dimer
> (from 5 to 2).  Does that tell me that I'm completely off?
> 
> Next I was thinking twinning but there is no indication for that (Yeates,
> sfcheck).
> 
> Currently I'm running Molrep with the dyad option and half of the
> suggested dimer (i.e. the initial monomeric model).  In C222 and C222_1
> Molrep comes up with a solution containing four models (as expected from
> Matthews coefficient) and good contrast values (11 and 120 for C222;  92
> and 85 for C222_1).  Looking at the solutions, I don't see dimers.  The
> other space groups are work in progress...
> 
> Furthermore, I plan to do the dyad search with a fixed model running
> Molrep 26 times with all solutions from Phaser.
> 
> 
> Am I chasing waterfalls, or does it sound like I'm up to something?  Any
> comments or suggestions are highly appreciated.  And, if you got that far,
> thanks for reading through all this ;-)
> 
> 
> 
> Andreas
> 
> 
> 
> And yes, I have whipped myself and a student for the last six months
> trying to get better crystals, derivatives, data, etc.
> 
> 
> 
> 
> Indexing in different space groups - output from XDS for signal/noise >=
> 0, all data:
> 
>         RESOLUTION     NUMBER OF REFLECTIONS    COMPLETENESS R-FACTOR
>           LIMIT     OBSERVED  UNIQUE  POSSIBLE     OF DATA   observed
> 
> P1 a      total      105246   62621    108903       57.5%       8.2%
> P1 b      total      105224   62606    108903       57.5%       8.2%
> P2        total      106172   50601     56092       90.2%       9.0%
> P2_1      total      106164   50594     56081       90.2%       9.0%
> C2 a      total      108603   38721     55096       70.3%      10.4%
> C2 b      total      107130   48446     55231       87.7%       9.2%
> C222_1    total      110472   27435     28860       95.1%      11.1%
> C222      total      110479   27441     28871       95.0%      11.1%
> 
>          R-FACTOR COMPARED I/SIGMA   R-meas  Rmrgd-F  S_norm/
>           expected                                     S_ano
> 
> P1 a       8.3%    85250    7.52    11.7%    26.4%    0.00
> P1 b       8.4%    85236    7.51    11.7%    26.4%    0.00
> P2         9.1%   102652    8.35    12.3%    26.6%    1.12
> P2_1       9.1%   102650    8.35    12.3%    26.6%    1.12
> C2 a      10.6%   102377    8.75    12.5%    23.5%    1.06
> C2 b       9.3%   101481    8.34    12.2%    25.4%    1.08
> C222_1    11.4%   110384   10.50    12.9%    19.9%    1.02
> C222      11.4%   110386   10.50    12.9%    19.9%    1.02
> 
> 
> 
> 

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