CCP4 bulletin board <[email protected]> wrote on 05/10/2007 09:31:59
PM:

> Hello,
> I'm an undergraduate and recently crystallized and obtained 2.9A
> diffraction data for a protein which is predicted to fold into a WD40
> 7-bladed beta-propeller structure (which has been crudely verified by
> cryo-EM by another lab). The space group appears to be I4(1) with
> unit cell parameters 118.936   118.936    85.456    90.000
> 90.000    90.000. Using a number of different search models (which I
> trimmed and aligned to my protein's sequence using Chainsaw) I have
> obtained a number of MR solutions placing 2 molecules in the AU with
> Phaser with high Z-scores (ranging from Z=9 to 12) that seem to pack
> together nicely, so I was hoping to use this technique to solve my
> structure. However, the initial Rfree for my best solution is
> relatively high (0.49) and all attempts to refine the structure
> result in the Rfree blowing up almost immediately. This makes me
> worry that the maps I'm generating may be too model-biased to use to
> generate a solution. I've tried using Prime and Switch to remove
> model bias but the resulting map looks worse than the starting map.
> As the predicted structure possesses so much radial symmetry (7-
> fold), I'm worried that my MR solutions will never be oriented
> correctly enough for me to be able to build a model. If anyone has
> any suggestions for tackling this kind of molecular replacement woe,
> I would greatly appreciate it. Otherwise I guess I'll just plan to
> collect experimental phasing information sometime in the near future.
>
> I'm not sure if this is the right place to be asking this question,
> perhaps you guys could direct me elsewhere.
>
> Thanks!
> -Scott


Hi Scott -

You can probably overcome the 7-fold symmetry problem manually.  I suggest
the following:

- Take a few of your search models that give the best solutions (are all
the solutions roughly superimposed?)

- Generate copies of the search model in each of the 7 orientations
corresponding to the 7-fold symmetry.  You might be able to generate a
symmetry operator to do this, but I'd probably just do it by eye using O or
something similar.

- Submit the rotated models to optimization using Phaser.  You'll use the
"Refinement and phasing" option in the GUI, or the "MODE MR_RNP" keyword in
the script file.  You'll feed Phaser a fake solution file (the .sol file)
with all Euler angles and fractional translations set to zero.

- Compare the resulting LLGs of the seven symmetry-related models.  With
luck, one of them will stand out from the others.  (Z-scores may not be
meaningful here - I forget whether Phaser calculates Z-score using a random
set of orientations, or just takes the mean and std. deviation of the
ensemble of solutions.)

- Take the best orientation (or all of them if one doesn't stand out) and
try Refmac refinement.  Again, one will hopefully look better than the
others.

Hope this works for you - it's certainly something to try while you're
trying to get derivatives...
And feel free to contact me if this wasn't clear enough for you to try it.

- Matt

--
Matthew Franklin , Ph.D.
Senior Scientist, ImClone Systems
180 Varick Street, 6th floor
New York, NY 10014
phone:(917)606-4116   fax:(212)645-2054


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