Dear Stefan,

Congratulations! It seems that you not only have pseudo-translation and pseudo 
4-fold symmetry in your diffraction, but also pseudo 8-fold symmetry in your 
protein molecules! An excellent case to learn a lot about space groups and NCS 
symmetry.

3) you are right, you might see pseudo-translation due to the internal 
pseudo-symmetry of your molecules. Does this fit with the molecular replacement 
solution you got?

5) Nothing is easier! Here the Refmac people really did an excellent job! Just 
give your P1 data set and P1 MR solution to Refmac and give the TWIN keyword. 
Refmac will sort everything out for you. I am currently working on a crystal 
form which is P43212, but where in most crystals the layers occasionally shift 
by half an unit cell resulting in a spacegroup of 1/2a, 1/2b, c with the same 
P43212 symmetry and 1/4 molecule in the unit cell. In a first round, I just 
processed in P1 and ran MR in P1 and found 1 dimer. Refmac detected and refined 
8 twin domains without any problems. Of course the 2-fold symmetry, generated 
by the dimer, is not twinning, so I still have to reprocess the data in P2 and 
refine with tetartohedral twinning.

I suspect, you might have something similar: perfect 4- or 4x-fold symmetry, 
but subsequent layers, or twin domains, are randomly shifted somewhat 
(left-right-up-down) and your NCS 4-folds to not coincide with the positions 
where crystallographic 4-folds should be. What kind of R-factors do you get 
when you refine your MR solution without twinning?

Herman


Von: CCP4 bulletin board [mailto:[email protected]] Im Auftrag von Stefan 
Gajewski
Gesendet: Dienstag, 22. Oktober 2013 06:29
An: [email protected]
Betreff: Re: [ccp4bb] Triclinic solution for a dataset with 
pseudo-translational symmetry, possible pseudo-centering and possible 
pseudo-merohedral twinning. How to proceed?

Dear Herman,
1) R-merge is about 11% in 4/mmm and only slightly lower in lower symmetry. 4/m 
would be an option since it has only four molecules in the cell but I do not 
get a solution for any group higher than P1.
2) Agreed.
3) Well, actually the pseudo 4-fold "works" when the pseudo translation also 
has a rotational component. I attached a diagram for the packing in the cell. 
My protein has a high internal pseudo-symmetry and overlaps pretty well with 
itself by rotation.

4) Each of the four proteins has a local 8 fold pseudo-symmetry of monomers.
5) tetartohedral twinning? what refinement software is able to deal with that?
Stefan



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