As Reza already pointed out, ITC cannot tell you anything about the 
sub-processes that underlay an equilibrium, be it complicated like 2A+B2 <-> 
AB2 + A <-> A2B2, or with (virtually) no intermediate 2A+B2 <-> A2B2 due to a 
fast second forward reaction , or a simple one-to-one model A+B2 <-> AB2.

Given the lack of additional information, its probably good to assume a simple 
one-to-one model and titrate either partner to the other.

If you have two separate binding steps (with similar Kd for B2 for A as well as 
AB2 for A), you would measure an apparent affinity and would see the 
stochiometrics according to  the inflection point (be it around equimolar 
excess or at 0.5 or 2, depending on whether you titrate A or B). If the 
reaction is more complicated and the the affinities for B2 for A differ 
significantly much from the affinity of AB2 for A, then a simple one-to-one 
would leave some notable information in the residual standard deviations 
(meaning, the residuals would not spread normally around the Regression line, 
but should show a wavy pattern).

Sorry for the long mail..

Matthias


Dr. Matthias Barone

AG Kuehne, Rational Drug Design

Leibniz-Forschungsinstitut für Molekulare Pharmakologie (FMP)
Robert-Rössle-Strasse 10
13125 Berlin

Germany
Phone: +49 (0)30 94793-284

________________________________
From: CCP4 bulletin board <[email protected]> on behalf of Michael Fairhead 
<[email protected]>
Sent: Thursday, October 3, 2019 5:59:22 PM
To: [email protected]
Subject: Re: [ccp4bb] ITC question -dimer vs monomer

Hello,
If you Google Alan cooper ITC insulin, you should find his work describing the 
study of insulin dimer:monomer equilibrium and the effect of cyclodextrin 
studied via ITC. This may be of some help.
Cheers
Mike

https://www.google.com/url?sa=t&source=web&rct=j&url=http://www.chem.gla.ac.uk/staff/alanc/itcdil.pdf&ved=2ahUKEwiOpoeisYDlAhUDThUIHX5mAyMQFjAAegQIAhAB&usg=AOvVaw0Ng-W03upy4DG12unFGDY3


From: CCP4 bulletin board <[email protected]> On Behalf Of Bernhard Rupp
Sent: 03 October 2019 16:52
To: [email protected]
Subject: Re: [ccp4bb] ITC question -dimer vs monomer

I am not looking for anything yet – I wonder what – if any – the consequences 
of doing it one way or the other would be.
I am reasonably certain that any difference affects the analysis.

Thx, BR

From: CCP4 bulletin board <[email protected]<mailto:[email protected]>> 
On Behalf Of Keller, Jacob
Sent: Thursday, October 3, 2019 17:41
To: [email protected]<mailto:[email protected]>
Subject: Re: [ccp4bb] ITC question -dimer vs monomer

I don’t understand what you are trying to do—are you trying to show, by the 
difference in ITC response, that the predictions you made about the 
oligomerization are true?

JPK

+++++++++++++++++++++++++++++++++++++++++++++++++
Jacob Pearson Keller
Research Scientist / Looger Lab
HHMI Janelia Research Campus
19700 Helix Dr, Ashburn, VA 20147
Desk: (571)209-4000 x3159
Cell: (301)592-7004
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From: CCP4 bulletin board <[email protected]<mailto:[email protected]>> 
On Behalf Of Bernhard Rupp
Sent: Thursday, October 3, 2019 11:06 AM
To: [email protected]<mailto:[email protected]>
Subject: [ccp4bb] ITC question -dimer vs monomer

Hi Fellows,

please let me ask the respective experts an ITC question: I have 2 proteins, 
stable and dialyzed in identical buffer.
A is a monomer and B an obligate dimer. I suspect that eventually a A2B2 dimer 
will form.

Intuitively, it should make a difference whether I titrate the dimer with the 
monomer or vice versa.
In the first case, a momomer would initially meet a lot of free dimers, and I 
would expect that randomly, a AB2 complex
is more likely to form than a A2B2 (let’s disregard any more complex 
colligative/cooperative effects).

If I drip the dimer into the monomer pool, it is quite likely that the B dimer 
meets 2 free As, and I get right away a higher population of A2B2s.

Maybe at dilutions of ITC and with sufficient equilibration that is not an 
issue at all (again, absent any cooperative effects that might alter the first 
Kd vs. the second, despite the sites on the dimer are at least initially 
equivalent).

Can someone guide me towards literature about this or perhaps share some 
first-hand experience?

Many thanks, BR

------------------------------------------------------
Bernhard Rupp
http://www.hofkristallamt.org/<https://urldefense.com/v3/__http:/www.hofkristallamt.org/__;!oCotSwSxbw8!SE9mT6grUy1tHFSKLCraXt4bhlDri03OEMEyqQUCLAVSLsg3vwn0GTQtxbStgtNvxBs$>
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+1 925 209 7429
+43 676 571 0536
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Many plausible ideas vanish
at the presence of thought
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