Hello,

It’s also amazing how one can mess up bond lengths so badly without actually 
damaging the angles—very interesting!

In this instance, it looks like an amyloid fibril, so each peptide is likely 
nearly perfectly in one plane. This means that any problem would leave the 
dihedral angles fine (consistent with the perfect Ramachandran distribution 
shown). If the model were a bit stretched in all directions, most bonds would 
be too long but very few of the bond angles would be distorted (also consistent 
with the low number of bond angle outliers shown).

Then, one reason why the model would be stretched in all directions could be 
that the pixel size of the cryoEM map is inaccurate (a bit larger than 
expected).

It seems like you have already identified the problem with the amide cap, but 
it might be a good idea to check the pixel size calibration just in case.

I hope this helps,

Guillaume


On 20 Feb 2025, at 02:50, Das, Abhinaba 
<[email protected]<mailto:[email protected]>> wrote:

Hi Kasper,

Yes, you are correct. There are 80 chains with 480 residues. Each of these 80 
chains has a C-amide capping. It’s hexapeptide whose sequence we know and 
matches the density. Unlikely they need to be merged. More precisely, the 
number of NH2 present equals to number of bonds of unrealistic length. Somehow 
the program thinks that the ideal value for the terminal C-N linkage should be 
2.329A but it’s an amide bond.

Best,
Abhi
From: Kaspar Hollenstein 
<[email protected]<mailto:[email protected]>>
Date: Wednesday, February 19, 2025 at 7:47 PM
To: Das, Abhinaba <[email protected]<mailto:[email protected]>>
Cc: [email protected]<mailto:[email protected]> 
<[email protected]<mailto:[email protected]>>
Subject: Re: [ccp4bb] [External] Re: [ccp4bb] Low atom inclusion score error 
while depositing to PDB

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Hi Abhinaba,

Based on your screenshot of the validation summary, your model contains 480 
residues spread over 80 chains. Does that match what is in your sample? Is it 
possible that some or all of these amino acids need to be merged into larger 
polypeptide chain(s)? Perhaps it is not a coincidence that the number of chains 
is equal to the number of bonds of unrealistic length.

Best,
Kaspar

On Feb 19, 2025, at 2:36 PM, Das, Abhinaba 
<[email protected]<mailto:[email protected]>> wrote:

Thanks Phil and Pavel for pointing it out. The error is likely (I think) of the 
NH2 ligands not being detected as proper amide linkage.

<image002.png>

<image001.png>

From: Pavel Afonine <[email protected]<mailto:[email protected]>>
Date: Wednesday, February 19, 2025 at 2:17 PM
To: Das, Abhinaba <[email protected]<mailto:[email protected]>>
Cc: [email protected]<mailto:[email protected]> 
<[email protected]<mailto:[email protected]>>
Subject: [External] Re: [ccp4bb] Low atom inclusion score error while 
depositing to PDB
Hi Abhinaba,

Here is my interpretation of this…

The geometry statistics are overall fine, except for the impossibly bad bond 
rmsd (overall, and greater than 4σ). This can't be true for an okay model. So 
I'd stop deposition and check/fix that first! It’s also amazing how one can 
mess up bond lengths so badly without actually damaging the angles—very 
interesting!

Overall, CCmask suggests the model fits the map okay as a whole. To get a 
better local picture, you’d need to check values per chain and residue.

I wouldn’t worry about the atom inclusion score at all—it’s a pretty 
meaningless quality metric, and I could spend an hour or two explaining why.

Bottom line: Based on the information you’ve supplied, your biggest concern 
should be bad covalent bonds.

Good luck!
Pavel

On Wed, Feb 19, 2025 at 10:42 AM Das, Abhinaba 
<[email protected]<mailto:[email protected]>> wrote:
Dear Community,

I'm trying to deposit an EM volume map and the corresponding model to the PDB. 
Though validation in Phenix doesn't show anything wrong, the deposition 
validation indicates a low atom inclusion score. Not sure why the error… Thanks 
in advance for any insights.


<image001.png>

<image002.png>
<image003.png>

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