Dear Julia,

Thanks a lot for your input. I looked at your pdb/maps and yes, the context
is similar, with the M2+ ion sitting near a His side chain, without further
ligands nearby. Although this is not canonical chemistry for ions like
these, I agree it must be some sort of water-metal ion adduct where waters
might not be perfectly localized.

Cheers
Talis

On Mon, Mar 9, 2026 at 12:00 PM Julia Griese <
[email protected]> wrote:

> Hi Talis,
>
>
>
> I agree with everyone else, this has to be a metal ion. It is not strange
> to see something like this, a metal ion with only one coordinating residue
> on the protein surface, in a crystal that has been grown or soaked in a
> high concentration of metal ions. A site like that is of course extremely
> unlikely to be occupied in solution or to be physiologically relevant.
>
>
>
> I would also agree with Jon that it is most likely to be Cd because that
> would be the strongest binder, but it depends on the relative amount of
> each metal ion in your mixture. (See Irving-Williams series.) Since they’re
> all at 5 mM in your case, it should be Cd. It will have loosely bound water
> filling up the remaining coordination sites, hence the disk-like appearance
> of the density.
>
>
>
> I have had a site exactly like that in a protein that I’ve worked with,
> see e.g. PDB 4HR0, Mn(II) bound to His130. The site follows the
> Irving-Williams series exactly. We dissected the origins of metal
> specificity of the heterodinuclear Mn/Fe site of this protein, so I spent a
> lot of time soaking the crystals with mixtures of different metal ions and
> figuring out which ends up where using anomalous diffraction, therefore I
> know exactly what is bound to all the sites, including the His130 site,
> under which condition. Although not physiologically relevant, for me this
> site was useful as an internal reference to distinguish good from bad
> datasets. It is almost certainly overkill for you to go to that amount of
> effort because it’s not a physiologically relevant site. I would go with
> Cd.
>
>
>
> Best,
>
>
>
> Julia
>
>
>
> --
>
> Dr. Julia Griese
>
> Associate Professor (Docent)
>
> Principal Investigator
>
> Department of Cell and Molecular Biology
>
> Uppsala University
>
> BMC, Box 596
>
> SE-75124 Uppsala
>
> Sweden
>
>
>
> email: [email protected]
>
> phone: +46-(0)18-471 4982
>
> Griese lab
> <https://www.uu.se/en/department/cell-and-molecular-biology/research/structural-biology/griese-lab>
>
>
>
>
>
> *From: *CCP4 bulletin board <[email protected]> on behalf of Eleanor
> Dodson <[email protected]>
> *Date: *Monday, 9 March 2026 at 07:43
> *To: *[email protected] <[email protected]>
> *Subject: *Re: [ccp4bb] Unusual density around His
>
> Anomalous peak height can give a clue.. any S atoms should also be marked
> in the anom diff map and the relative peak heights might help decide?
>
>
>
> On Sun, 8 Mar 2026 at 22:39, Jon Cooper <
> [email protected]> wrote:
>
> It must be cadmium which has atomic number 48 versus 27 for cobalt and 28
> for nickel. I would check that it could form a stoichiometric complex with
> the protein at 5 mM with whatever protein concentration you used. It almost
> certainly could.
>
> Best wishes, *Jon Cooper* (Emeritus at UCL) [email protected]
>
> Erratum and other hopefully useful things: https://crxp.org.uk
>
>
>
> Sent from Proton Mail <https://proton.me/mail/home> for Android.
>
>
>
> -------- Original Message --------
>
>
> On Sunday, 03/08/26 at 10:48 Greenstone talis <
> [email protected]> wrote:
>
> Dear all,
>
> I have came across an unusual density feature in a structure refined at
> 1.65 Å and would appreciate some opinions.
>
> There is a strong Fo–Fc positive density feature around the imidazole ring
> of a His residue. The density looks broad and somewhat flat, almost like a
> disk around the ring, rather than a compact peak.I initially thought it
> might be a metal because the crystallization cocktail contains Cd²⁺, Mg²⁺,
> Ni²⁺ and Co²⁺. Besides Fo-Fc results with one or another option, which of
> course is relevant, what still confuses me is the absence of coordinating
> ligands nearby. The density appears partially fused to the His side chain
> rather than sitting clearly next to it. I am attaching three views of the
> density from different angles (2Fo–Fc at 1 sigma, Fo–Fc at 3 sigma).
>
> Does this look like something people have seen before? Could it indicate
> some PTM (for example oxidation or another chemical change), or is there
> another explanation that I should consider?
>
> Any suggestions would be very helpful.
>
> (PROTEIN SOLUTION: TRIS AND NaCl)
>
> (CRYSTALLIZATION COCKTAIL: 0.005 M MgCl + 0.005 M NiCl + 0.005 M CoCl +
> 0.005 M CdCl + 0.1 M HEPES pH 7.0, PEG 3350 + ETHYLENE GLYCOL)
>
> Thanks in advance.
>
> Talis
>
>
>
>
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