Hi,

I have a very peculiar crystal structure which has been challenging to solve:

Prot-X + VHH – nominally expected to be a 1:1 complex

Data auto-process as C2
CRYST1  132.084   56.245   86.262  90.00 108.40  90.00 C 1 2 1

2 copies of Prot-X easily located by molecular replacement, Phaser fails to 
find the VHH

Manual assessment of density in crystal voids reveals a density patch about the 
right size / shape for a VHH.
Manual placement of a VHH into this density shows it lies across the C2 2-fold 
so the lattice symmetry places an inverted copy of the placed VHH in the same 
location.

Expanding the symmetry to P1 gives 4-chains, arranged in a square ring, with a 
hole in the middle where the VHH sits, sterically only 1 VHH can sit in the hole
CRYST1   56.245   71.780   86.262 106.88  90.00 113.07 P 1

I overlaid a computational model of Prot-X + VHH onto each of the 4 chains, 2 
positions align reasonably with the density (related by 2-fold as expected), 
the other 2 do not.
If I refine either of these models in P1 with twin refinement then I get a 
reasonable map, especially where the VHH contacts Prot-X as the alternative 
2-fold related position does not overlap at the contact surface.

Some parts of the density look really good, some are a bit confused / look like 
some rebuilding is required, though this may be largely residual density from 
the alternate orientation.

Data resolution is modest with fairly noisy outer shell data so I can process 
to 2.6A with much better stats in C2 but only 2.8A and significantly worse 
stats in P1.

I have had similar cases before in monoclinic/orthorhombic and 
trigonal/hexagonal and was able to obtain better maps by symmetry expanding the 
higher resolution processing from the higher symmetry spacegroup to the lower 
symmetry spacegroup rather than processing in the lower symmetry.

I would therefore like to take the C2 processed data, symmetry expand to P1 and 
then use these expanded data for twin refinement to try and improve map quality.

I am however not sure how to expand / transform the data from the C2 lattice to 
the P1 case.

Help appreciated…

Thanks, Charlie.

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