Hi all:

Cordilleran Flycatcher is a widespread and common breeding species in 
Colorado's foothills and mountains, a species that is very familiar to many of 
us, particularly to those hosting breeding birds on their houses or 
outbuildings.  However, I believe that that familiarity and abundance has made 
for overcertainty in reporting of the species by Colorado's birders.  While the 
proximal impetus for this essay was from my task as a reviewer of Colorado 
eBird records, it is a subject that I have pondered for quite some time.  In 
fact, I have posted to this venue previously on the topic.

Publications in the 1980s and beyond established the case for splitting what 
was then known as Western Flycatcher into two component species, Cordilleran 
and Pacific-slope flycatchers. While the breeding ranges of the two are well 
established and involve a bit of geographic overlap, nearly all else that we 
know about the two species' distributions is not supported all that well with 
hard data. The reason for this is that the two species are VERY difficult to 
differentiate in the field AND in the hand.  Rocky Mountain Bird Observatory 
has spent years catching migrants on the plains and most such birds have been 
reported as Western Flycatcher, because it is so very difficult to be sure of 
what one has in hand. Yes, there are known differences. The Male Position Note 
differs noticeably between the two species (though with some apparent hybrids 
or individuals giving both calls or intermediate calls in the overlap zone [and 
elsewhere?]). However, all other vocalizations overlap significantly, negating 
their use in definitive species identification. While there are some 
plumage-color differences, these are subtle and require extensive experience 
with both species in a variety of lighting conditions to use and, even then, 
are fraught with difficulty. Thus, identifying any single non-Male Position 
Note-calling Western Flycatcher to species is, essentially, a case of guessing. 
 In Colorado, I am quite comfortable with identifying all spring-migrant and 
breeding-season Western Flycatcher to species, specifically Cordilleran 
Flycatcher, though, if someone came up with data supporting the occurrence of 
Pacific-slope in the state during that time period, I would gladly reassess 
that opinion.  I believe that such IDs are "safe" because there are no data 
suggesting the occurrence of the other species in the state then.  Granted, 
that's a bit of circular thinking, but, hey, I'm only human.  

However, I do object to willy-nilly identifying fall birds as Cordilleran, 
particularly on the eastern plains.  That is because nearly across the board, 
Rocky Mountain breeding-bird species seem to be able to go north and south on 
their migrations without drifting off the the cordillera in any significant 
numbers, particularly in fall.  As example, Cassin's Vireo (a northwest-Pacific 
breeder) is much more common on Colorado's eastern plains in fall than is 
Plumbeous Vireo (which is rarish there then), which breeds within sight of many 
well-birded eastern-plains locations.  Cassin's Vireo is also not at all rare 
in the mountains and on the West Slope, though greatly outnumbered by Plumbeous 
Vireos there.  The comparison between the migrant Townsend's Warbler versus the 
breeder Black-throated Gray Warbler is similar, but on an even larger numerical 
scale.  Likewise, White-throated Swift, Red-naped Sapsucker, Purple Martin, 
Violet-green Swallow, Western Bluebird, western Veery, Virginia's Warbler, Sage 
Sparrow, Slate-colored Fox Sparrow, Mountain White-crowned Sparrow, and 
Gray-headed Junco are all, in fall, fairly rare to exceedingly so in eastern 
Colorado away from montane or foothill habitats.  Yes, there are bird species 
breeding in montane Colorado that are not rare on the eastern plains in fall, 
but most of those are more-widespread breeding species in which more-far-flung 
popluations might be masking the occurrence of actual Colorado breeders on the 
plains.

So, why is it that we willy-nilly report all Western Flycatchers in Colorado in 
fall as Cordilleran?  Because Pacific-slope has not been accepted as occurring 
in the state?  Forgive me, but that is a specious and even-more-circular 
argument.  If we don't figure out how to identify, or even consider, 
Pacific-slope Flycatcher, how can the species ever be accepted to the state 
list.  Again, consider how relatively common Cassin's Vireo and Townsend's 
Warbler are in the state in fall, both species that share breeding range with 
Pacific-slope Flycatcher.

Sincerely,

Tony Leukering
Villas, NJ

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