I remember reading somewhere in the SMTP RFC about the fact that e-mail addresses are to be case-insensitive.
domains yes, but everything before the @ can be case sensitive. Courier is case sensitive unless you tell it to not be like so:
touch /etc/courier/locallowercase
If you do that, courier will drop all incoming messages to lower case so you need to make sure that none of your accounts have upper case letters in them, or else you'll get a 550 again.
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