I think this bug may have been fixed upstream a couple of weeks ago when they updated the code to use GtkComboBox instead. See <http://svn.gnome.org/viewvc/evolution/trunk/composer/e-msg-composer-hdrs.c?revision=34620&view=markup> and note this callback:
g_signal_connect_swapped (
widget, "changed",
G_CALLBACK (signature_changed), hdrs);
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