I am running Gwibber 2.91.2. Epiphany also opens instead of Firefox when I click on links in Evolution; when I tried to click on the mailto:[email protected] link in Evolution, it said "Could not open link. The specified location is not supported." I don't know if that's the same problem or not. I have tried /opt/firefox/firefox (the actual location of my Firefox 4 Beta), /usr/bin/firefox (I linked /opt/firefox/firefox there in hopes that it might help), and selection "Firefox" from the dropdown menu in System->Preferences->Preferred Application. I also have Evolution listed as the preferred mail client.
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