And here's the original file.

evince prints it ok, FWIW (except for bug #120861, but that can be
worked around).

** Attachment added: "Pybus_et_al_2000_Genetics.pdf"
   http://launchpadlibrarian.net/8561964/Pybus_et_al_2000_Genetics.pdf

-- 
strange font behaviour - connected letters
https://bugs.launchpad.net/bugs/128074
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