I have the same issue, on Ubuntu 17.10. I solved it by moving the
download-to directory to home. My "Desktop" is a symlink to
DropBox/Desktop, which is a DropBox, with that may entail.

In summary - my suspicion is that it is DropBox to blame.

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You received this bug notification because you are a member of Desktop
Packages, which is subscribed to transmission in Ubuntu.
https://bugs.launchpad.net/bugs/700214

Title:
  File went missing as soon as download completed

Status in transmission package in Ubuntu:
  Expired

Bug description:
  Binary package hint: transmission

  I downloaded a file from the torrent
  http://releases.ubuntu.com/10.10/ubuntu-10.10-desktop-i386.iso.torrent.
  I saved the file to my Desktop. After the download completed,
  Transmission started seeding the file (I don't know if it actually
  uploaded any data). I opened the Desktop in Nautilus and the file was
  not there. I went back to Transmission (never having closed the app)
  and asked it to verify the download. The file immediately changed to
  red and said "Error: Can't find local data. Try "Set Location" to find
  it, or restart the torrent to re-download."

  Needless to say, this was quite frustrating. This happened within
  seconds of the download completing and I did no other activities which
  could have caused the file to be deleted or moved.

  Transmission version 2.04-0ubuntu2.

  ProblemType: Bug
  DistroRelease: Ubuntu 10.10
  Package: transmission-gtk 2.04-0ubuntu2
  ProcVersionSignature: Ubuntu 2.6.35-24.42-generic 2.6.35.8
  Uname: Linux 2.6.35-24-generic i686
  NonfreeKernelModules: nvidia
  Architecture: i386
  Date: Sat Jan  8 16:23:30 2011
  EcryptfsInUse: Yes
  ExecutablePath: /usr/bin/transmission
  InstallationMedia: Ubuntu 10.04 LTS "Lucid Lynx" - Release i386 (20100429)
  ProcEnviron:
   PATH=(custom, user)
   LANG=en_AU.utf8
   SHELL=/bin/bash
  SourcePackage: transmission
  XsessionErrors:
   (polkit-gnome-authentication-agent-1:1998): GLib-CRITICAL **: 
g_once_init_leave: assertion `initialization_value != 0' failed
   (gnome-panel:1995): GConf-WARNING **: Directory 
`/apps/panel/toplevels/bottom_panel_screen1/screen' was not being monitored by 
GConfClient 0xa1331a0
   (gnome-panel:1995): GConf-WARNING **: Directory 
`/apps/panel/toplevels/top_panel_screen1/screen' was not being monitored by 
GConfClient 0xa1331a0

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