It looks like the mailinglist server IPv6 address has been blacklisted:

https://check.spamhaus.org/listed/?searchterm=2a04:3545:1000:720:acc1:5bff:fe5e:459

The 'Show details' shows the entire 2a04:3545:1000:720::/64 network is blacklisted. Whois tells this is a network used by UpCloud Cloud Servers. So either the mailinglistserver or a server it shares its network with is compromised. If UpCloud gives its users their own /64, it would probably be the former. Otherwise it might make sense to ask them for one, as sharing IPv6 networks between customers is a no-no in the IPv6 world.

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