Norris,

I think you have a good point to illustrate the problem:

> Imagine the case where you have two phenotypically identical 
> populations that have different underlying genetics. If these 
> populations have different heritabilites, application of the identical 
> selective pressures could lead to dramatically different outcomes. In 
> this scenario only the responses to selective pressures would differ. 
> It would seem inconsistent to me to retro-actively claim that natural 
> selection was only operating in the one case where there was a 
> response. Rather, natural selection was only effective in producing 
> evolutionarily relevant change in one case.
There is no inconsistency, because for natural selection to act, the 
phenotype must have a connection (heritability) with its underlying 
genotype.  So, in your example above, let's just say that one 
population's phenotype was totally environmental (a good year, perhaps) 
while the other population's identical phenotype was genetic.  Well, in 
both cases perhaps the individuals with the same, "high quality" 
phenotype would be favored (leaving more descendants) but only the 
population with a genetic basis would leave behind the "tendency" that 
was based on the phenotypic expression of the genotype.  The other, in a 
different environment, would no longer show the same phenotype.

Did I explain well?

Cheers,

Jim

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