Increasingly, the influence of landcover (more commonly referred to as the
matrix) is being studied w.r.t species diversity and composition of forest
patches (or other habitat types of focal interest) embedded in those
landscapes.  Recently, this was a subject of discussion in one my lab
meetings based on the following article:

Conservation biology [0888-8892] Bhagwat yr:2005 vol:19 iss:6 pg:1853 -1862

In brief, the authors were comparing diversity of sacred groves in India
(range of patch sizes) at various distances from a large forest reserve. 
The landscape between the sacred groves and the reserve was predominantly
coffee estates with tree cover.  The authors found little to weak evidence
of either patch size effects or distance from the reserve on mean species
richness within the groves.  In conclusion, the authors stated that
biodiversity within the sacred groves was (positively) influenced by tree
cover the matrix.

If this study were testing the effect of matrix quality on biodiversity
within these patches however, would one not need to include a landscape were
the habitat patches (in this case being the sacred groves) were surrounded
land-uses without tree cover?  This has very important implications for the
design of field surveys to assess matrix effects on biodiversity.

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