I believe I remember hearing a story of a pine that wouldn't grow in Pacifi=
c islands until its mycorrhizal symbiont was introduced though I couldn't f=
ind any publications about that specific case. I'd like to hear more about =
this topic.
=20
Here's some information from the U.N.'s website:=20
=20
FOOD AND AGRICULTURE ORGANIZATION OF THE UNITED NATIONS
http://www.fao.org/docrep/006/j1467e/J1467E06.htm#TopOfPage
=20
...care should be exercised over the introduction of exotic mycorrhizae to =
promote the growth of forestry trees. In general, these introductions may b=
e expected to be beneficial, but it is extremely difficult to anticipate or=
 assess the impact of such species on an ecosystem (Case Study 19). Introdu=
ctions of these species should therefore be approached with caution (partic=
ularly in the case of oceanic islands) and efforts made to monitor their su=
bsequent establishment, spread and ecological impact.
=20




Case Study 19: Possible impacts of mycorrhizal introductions associated wit=
h forestry trees=20
There is relatively little information on the impact of introduced mycorrhi=
zal fungi, except on their direct benefit to associated trees. Their possib=
le effect on the invasiveness of their associated trees (or other ecologica=
l effects) has not been widely investigated.
Introductions of symbiotic mycorrhizae may increase the invasiveness of exo=
tic plants, but at present there is little evidence of this. However, there=
 is clear evidence from Hawaii that fungal symbionts of alien tree species =
can significantly alter the natural ecosystem. In Hawaii, Myrica faya, an i=
nvasive alien tree, forms a symbiotic relationship with Frankia spp. (an ac=
tinomycetal genus that forms nitrogen-fixing root nodule symbioses). It is =
not known whether Frankia was naturally present in the soil in Hawaii, but =
the interaction between M. faya and Frankia significantly increases soil ni=
trogen levels. These high nitrogen levels, in turn, provide a nutrient-rich=
 substrate for other fast-growing alien species, which can outcompete slowe=
r-growing native species. The high-nitrogen soil also attracts worms (anoth=
er alien to Hawaii) that in turn attract pigs (also alien). The pigs distur=
b the soil looking for worms, creating germination sites for invasive alien=
 weed species. There is thus a whole series of negative impacts associated =
with this particular tree-mycorrhizal relationship.
In New Zealand, the introduced ectomycorrhizal fungus Amanita muscaria has =
become naturalized in some Nothofagus forests. This fungus was introduced t=
o New Zealand relatively recently, but before 1997 it had only been reporte=
d in three or four native forests within the Nelson Lakes National Park. Re=
cent records, however, suggest that it is now widespread in indigenous fore=
sts in the northern half of the South Island. In most localities where it i=
s found, Amanita muscaria is present at numerous separate sites. This distr=
ibution may be due to some local characteristic of the environment, the fun=
gus, or the host, or it may be the result of local spread following a singl=
e invasion. So far, it has been recorded only in association with Nothofagu=
s spp., with no reports of it occurring with the only other indigenous ecto=
mycorrhizal trees in New Zealand, Leptospermum scoparium or Kunzea ericoide=
s.
This system provides an ideal opportunity to research various questions rel=
ating to the effect of mycorrhizal species on the forest ecosystem, includi=
ng:


The effect of these exotic mycorrhizal associates on the indigenous Nothofa=
gus particularly with respect to nutrient uptake, resistance to root diseas=
es, etc.

The impact of these invasive fungi on the diversity of native ectomycorrhiz=
al fungi normally expected at the invaded sites.

The possible effect of these exotic mycorrhizal fungi on the invasion of No=
thofagus forests by exotic mycorrhizal trees. For example, in New Zealand, =
there is no evidence that exotic conifers can utilize indigenous mycorrhiza=
l fungi, and it could be that this restricts the spread of these conifers i=
nto New Zealand's native forests. However, since Amanita muscaria is often =
found as a mycorrhizal associate of introduced conifers (e.g. Pinus spp. an=
d Pseudotsuga menziesii), establishment of the fungus in native forests may=
 make it easier for Pinus spp. to invade them.=20
Sources: http://www.landcareresearch.co.nz/research/biosecurity/fungal/; Ca=
ndace J. Felling, personal communication, 2002; Tim Low, personal communica=
tion, 2002; Peter Johnston, personal communication, 2002. See also Lutzow-F=
elling et al. (1995); Vitousek (1986, 1992). > Date: Fri, 7 Dec 2007 13:00:=
22 -0500> From: [EMAIL PROTECTED]> Subject: Re: "Bio-dynamic" composting sta=
rters> To: [email protected]> > Dear Thomas,> > The potential for s=
oil invasions seems real. I have often wondered about the> generic "mycorrh=
izae in a can" being attractively marketed in organic> gardening catalogs. =
Terrestrial ecologists are still revealing so much about> this long underap=
preciated community. By the way, one cheap and local way to> introduce soil=
 biota into your compost pile is mixing some of your soil in> the pile.> > =
Scott> ---> > Scott Ruhren, Ph.D.> Senior Director of Conservation> Audubon=
 Society of Rhode Island> 12 Sanderson Road> Smithfield, RI 02917> > 401-94=
9-5454> > -----Original Message-----> From: Ecological Society of America: =
grants, jobs, news> [mailto:[EMAIL PROTECTED] On Behalf Of Thomas =
Hardy> Sent: Friday, December 07, 2007 12:01 PM> To: [EMAIL PROTECTED]
EDU> Subject: "Bio-dynamic" composting starters> > Does anyone know if use =
of commercial preparations of bio-dynamic compost> starters. chock-full of =
bacteria, soil fungi and perhaps a nematode or two,> if used in other regio=
ns of the world might raise questions of> "exotics-transfer"? Or are soil m=
icro-flora pretty much "homogenized" after> millions of years of climatic "=
adjustments", regardless of location?> > Thomas Hardy> > > ----------------=
-----------------> Looking for last minute shopping deals? Find them fast w=
ith Yahoo! Search.
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