Highly detailed models seldom work well. In a classic paper by Weigert he showed that the best models have an intermediate level of complexity beyond which performance gets worse. The interactions between closely related species are so difficult to quantify that some degree of aggregation is essential.

Galtman didn't specify what the purpose of the model was and what he expected it to do, but for a long time people have modelled predator-prey systems successfully. We did this by avoiding the use of parameters that we couldn't measure. Sure the models don't answer every question, but so long as they answer the questions we posed that was enough.

That doesn't mean that every question can be answered by modelling. Many years ago we had a joint workshop with the fishing industry and they wanted us to predict biomasses of cod and haddock. I pointed out that we had already developed a model that predicted the total landings of cod+haddock, but the two stocks varied unpredictably with inverse correlation so that we could not break the total down into the two components. They were dissatisfied with that, but that was the best we could do.

Bill Silvert


----- Original Message ----- From: "joseph gathman" <[email protected]>
To: <[email protected]>
Sent: Thursday, January 22, 2009 9:12 PM
Subject: [ECOLOG-L] Subject: Ecology Certainty Uncertainty Illusion/Delusion, was Re: Ecological Modelling


That means, I suspect, that ecology is, in the view of literal believers, > "doomed" to be "applied."

I don't know about that, but I DO think that what may well be doomed is our ability to accurately model complex systems. In my own limited attempt at modelling, I tried to model a simple predator-prey system, but it quickly led me into a rat's nest of uncertainties. It would have taken years to collect reliable data from field samples and experiments in order to make the model realistic. And that was with an artificially simplified model system. I can't imagine what it would have taken to build a "real" model.

Even the climate modellers acknowledge that over the last decade, all they've managed to do is confirm what their models CAN'T do (and likely never will). Somebody should tell the IPCC that bit about models not being predictive tools.

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