Dear all,

Currently the NCB Naturalis has a position available for a PhD student for a 
project on:

Evolution and biogeography of tropical understorey plants

Modern tropical rain forests are one of the most important and species rich 
biomes on the planet. 
They are generally characterized by a stratified closed canopy, which creates 
deep shade for 
understorey plants. Numerous flowering plant lineages show particular 
adaptations to grow in 
these low-light environments (e.g. leaf shape, dependence on mycorrhizal 
fungi). How this 
diversity of tropical understorey plants evolved remains largely unexplained. 
In this project, the 
candidate will use a combination of multi-gene phylogenetic analyses, molecular 
clock dating, 
diversification analyses, and biogeographical reconstructions to explore the 
relationship between 
rain forest history and shade tolerance in tropical flowering plants. The 
student will be expected to 
use the herbarium collections at NCB Naturalis and to take part in at least one 
collection trip to 
Southeast Asia. This project will require inferring phylogenetic relationships 
of particular lineages 
of tropical understorey plants, as well as constructing hypotheses of the 
biogeographical histories 
of these clades. The candidate will integrate methods in systematics, 
collection-based research, 
fieldwork, morphology, phylogenetics, and historical biogeography. The goal 
will be to try to 
discover common patterns in the evolution of rain forest understorey plants. 
The results will 
provide insights into the history of modern rain forests, and will be 
transferrable to conservation 
research.

Skills: The successful applicant will have proven knowledge of molecular 
phylogenetics and plant 
systematics.

General requirements
You have a master degree in systematics, evolutionary biology or other relevant 
disciplines. You 
have a scientific and critical attitude, excellent time management and 
organizational skills, the 
ability to work independently, good verbal and written communication skills. 
Fluency in English 
required.

We offer
A fulltime contract (36 hours per week) for a period of four years, starting 
June 1st 2011. However, 
the start date can be postponed by a few months if necessary. A salary that is 
comparable with 
salary for PhD’s at Dutch Universities. All our employees are incorporated into 
a pension fund. The 
successful candidate will be employed by NCB Naturalis in Leiden. The candidate 
will alsobe 
affiliated to the University of Leiden. The appointment must lead to the 
completion of a PhD thesis. 
During your appointment you will be supervised by Dr. Vincent Merckx.

Procedure
You are invited to submit your application including your curriculum vitae, a 
list of publications, up 
to five keywords describing your research interests, and the names and e-mail 
addresses of at 
least two persons that can be contacted for reference (and who have agreed to 
be contacted) 
before April 15th 2011 by e-mail to; [email protected], or by mail; 
NCB Naturalis, HR 
department, PO box 9517, 2300 RA Leiden, the Netherlands.
For more information on the current research and facilities, see the websites 
of the founding 
partners of the NCB Naturalis, on www.ncbnaturalis.nl - www.nhn.leidenuniv.nl - 
www.science.uva.nl/zma - www.bis.wur.nl/UK/Organisation/Herbarium

Any additional information can be requested from:
Prof. Dr. E.F. Smets, scientific director 
Netherlands Centre for Biodiversity Naturalis 
E-mail: [email protected] 
Phone: +31 71 5687 713

Or:
Dr. Vincent Merckx      
Research Fellow
Netherlands Centre for Biodiversity Naturalis (section NHN)
P.O. Box 9514, 2300RA Leiden, The Netherlands
email: [email protected]
phone: + 31 71 5273570
fax: + 31 71 5273522

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