Wayne, I think the notions that have been expressed on the list represent 
semantic differences rather than differences in understanding how roots 
function.  I think that those who have written accept the following:

Roots are hydrotropic, but the hydrotropism acts along a gradient, and acts 
over a very small distance.  Rather than individual roots turning or growing 
toward water at some distant location, they grow differentially in different 
moisture fields.  Whether one calls this behavior "seeking" or simply 
differential growth is dependent on one's application of the term to inanimate 
behavior or not.

I can no longer speak for what is being taught, but I always taught that roots 
exhibit positive gravitropism but negative phototropism, while the shoot 
exhibits the opposite, and described classic experiments (and in elementary 
courses had students carry out such experiments) that demonstrate such 
behavior.  Those tropisms do result in roots 'moving' toward moisture in soil.  
So far as hydrotropism, I did not teach it, because I did not recognize that 
the enhanced growth under higher moisture conditions compared to lower moisture 
conditions was a tropism.  I now know that hydrotropism is involved, though it 
is minor compared to the differential growth along a gradient.  I would teach 
that, were I still teaching.

mcneely

---- Wayne Tyson <[email protected]> wrote: 
> Honorable Ecolog Forum:
> 
> What does it matter, for the advancement of science and ecology in 
> particular, whether or not the root of the matter is resolved such that the 
> state of knowledge in this matter is articulated with clarity to the public 
> and those who inform the public (such as science writers, TV and movie 
> producers, reporters, etc.)?
> 
> There seems to be at least two schools of thought (or at least opinion) 
> concerning whether or not roots are genetically programmed to seek water.
> 
> What is actually being taught in the schools with respect to this issue? 
> What have you, as an ecologist, been taught and what do you teach (if you 
> teach)?
> 
> WT 

--
David McNeely

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