PhD Studentship available at the University of Stirling
"The extent and magnitude of negative density dependence among tropical
rainforest trees"

In addition to harboring two-thirds of terrestrial biodiversity and half of
terrestrial carbon, tropical forests provide a substantial fraction of
global ecosystem services, including the provision of timber, fuel, oxygen
and climate regulation. As anthropogenic threats to tropical forests
continue to mount, there is increasing need for ecosystem management that
can strike a balance between guarding against species extinctions and
sustainably providing ecosystem services. Effective management, in turn,
requires the ability to predict the dynamics of ecological communities into
the future, even into potentially novel environmental conditions. 

Species diversity and coexistence are stabilized by an inverse relationship
between population density and per-capita performance, measured as the vital
rates of growth, survival recruitment and reproduction. As such, negative
density dependence (NDD) has been a long-standing focus of ecological
research. Many studies have quantified the magnitude of NDD at single
ontogenetic stages. Even so, there is no synoptic understanding of the
importance of NDD across the life-cycle of any tropical tree. The objective
of this project is to determine the extent and magnitude of NDD for a
representative set of neotropical rain-forest trees. The successful
applicant will determine the degree to which each vital rate is affected by
population density, and at which ontogenetic stages. Achieving this
objective is key for predicting community dynamics, given the potent
diversity-stabilizing effect of negative density dependence. 

The successful applicant will combine existing data on the vital rates of
tropical rain forest trees with supplemental experimental and observational
data. These data will be used to model each vital rate as a function of
ontogeny and population density. As such, the ideal candidate will have
skills (or interest) in both field-based ecology and statistical analysis.
The project will be supervised by C. E. Timothy Paine, and will involve
close collaboration with project partners in the UK and French Guiana.
Depending on the applicantÂ’s skills and interests, the project may include
extended periods of field-based research in the forests of French Guiana
(South America). 

To apply, please send 1) a cover letter, 2) a curriculum vitae, 3) a copy of
University grades, and 4) contact details for two academic referees as a
single .pdf document to C. E. Timothy Paine ([email protected]) by
February 1, 2013. The entry qualification is a First Class or Upper Second
Class Honors degree and/or a relevant Masters degree in a biological or an
environmental science (or equivalent qualifications gained outside the UK).
Given the short duration of the PhD programs in the UK (3 years), preference
will be given to applicants with previous research experience or a MSc
degree. Experience in ecological fieldwork or statistical analysis, and
fluency in French, would be assets. Your cover letter should clearly set out
your suitability and motivation for this PhD, making reference to your past
experience and achievements. Please email me if you have further questions
about the studentship. 

Funding Note:
The studentship is part of a competitive NERC-quota PhD studentship awarded
to the Biological and Environmental Sciences department of the University of
Stirling. The top candidate will be put forward for a final round of
internal selection against candidates from other labs in the Department. The
studentship will cover tuition fees and provide a stipend for UK/EU
students. Overseas students are invited to apply, but must independently
obtain financial support for all non-UK fees.

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