6 PhD / postdoc positions available on eco-evolutionary assembly of 
ecological communities

At the Centre for Ecological and Evolutionary Studies of the University of 
Groningen in the Netherlands, there are 6 vacancies for PhD students and/or 
postdocs, as part of the research program “On the origin of species 
assemblages” which is funded by a VICI grant awarded to Prof. Rampal S. 
Etienne.

CONTENT

This program aims to develop a fully stochastic, dynamical, eco-evolutionary 
and data-friendly theory of community assembly, and testing and informing 
this theory with model-oriented experiments and field studies of both macro-
organisms and micro-organisms. 
The theory will contain models of speciation, extinction, immigration, and 
evolutionary change that vary in spatial, phylogenetic and biotic 
complexity, and will be designed for confrontation with data. The models 
include but are not limited to birth-death models, individual-based 
simulation models, and models of adaptive dynamics. New modelling directions 
are encouraged.
The evolutionary experiments will involve the mite Tetranychus urticae (in 
collaboration with Dries Bonte at the University of Ghent, Belgium) and the 
bacterium Escherichia coli (in collaboration with Joana Falcao Salles at the 
University of Groningen) which are ideal model organisms due to their short 
generation times. The experiments will provide insight into how diversity 
affects diversification, a great unknown in current macro-evolutionary 
theory. 
The theory will be applied to naturally occurring microlandsnails in South-
East Asia (in collaboration with Menno Schilthuizen at Naturalis 
Biodiversity Center, Leiden, The Netherlands), and micro-organisms in 
geothermal pools in New Zealand (in collaboration with Gavin Lear at the 
University of Auckland, New Zealand).
As a general outline, for the theoretical part, both potential PhD students 
and postdocs can apply. The mite and microsnail projects will each require a 
PhD student, the E. coli and geothermal pool projects together will require 
a postdoc. However, candidates who are willing and able to handle a 
combination of theoretical and empirical work are particularly encouraged to 
apply.

PROFILE OF CANDIDATE

The candidates should
- hold an MSc degree in ecology or evolutionary biology, or an MSc degree in 
mathematics or physics with experience/interest in applications to biology
- be willing to work in a multidisciplinary team
- have good communication skills in English (speaking and writing)

INFORMATION AND APPLICATION

More information can be requested from Prof. Rampal Etienne 
([email protected]). Applications can be sent to the same address and 
should contain:
- Motivation letter (max. one page)
- Up-to-date CV
- List of study grades
- Names of two references
Incomplete applications will not be considered. Applications will be 
considered until the positions are filled.

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