This year's winner is Alejandro Rico-Guevara from the University of Connecticut 
for his paper "Bills as daggers? A test for sexually dimorphic weapons in a 
lekking hummingbird" published in Behavioral Ecology, a close collaboration 
with Marcelo Araya-Salas, a graduate student at New Mexico State University.  
He conducted this research while working with Margaret Rubega. This 
revolutionary work demonstrates that Long-billed Hermit males use their bills 
as weapons in territorial defenses and offers a new explanation for sexual 
dimorphism in bills of hummingbirds. This 4 year study combined behavioral 
observations in the field with territory mapping, mark-recapture, ontogenetic 
changes, detailed morphological analysis, and performance experiments. 
Fieldwork was carried out at La Selva Biological Station, Costa Rica, which is 
operated by the Organization for Tropical Studies.
Runner-up is María-José Endara from the University of Utah for her paper 
"Divergent evolution in antiherbivore defenses within species complexes at a 
single Amazonian site" published in Journal of Ecology. She was co-advised by 
Phyllis Coley and Thomas Kursar and is an alum of 2 OTS courses. Her research 
addressed a challenging and important question examining how insect herbivores 
may drive the maintenance and origin of tropical tree diversity. She conducted 
several years of fieldwork in a remote site in the Amazon, and coupled 
fieldwork with lab work, including UPLC-mass spectrometry analyses of secondary 
metabolites and sequencing DNA of herbivores to reconstruct a phylogeny.
Three papers received Honorable Mention. Timothy (Trevor) Caughlin from the 
University of Florida for his paper "Loss of animal seed dispersal increases 
extinction risk in a tropical tree species due to pervasive negative density 
dependence across life stages" published in Proceedings of the Royal Society B: 
Biological Sciences.  He worked with Douglas Levey, who is now at the National 
Science Foundation. Luke Owen Frishkoff from Stanford University for his paper 
"Loss of avian phylogenetic diversity in Neotropical agricultural systems" 
published in Science, while working with Gretchen C. Daily. Alison Ravenscraft 
also from Stanford University for her paper "Nutrient acquisition across a 
dietary shift: Fruit feeding butterflies crave amino acids, nectivores seek 
salt" published in Oecologia. She conducted her research with co-advisors Carol 
Boggs, now at the University of South Carolina, and Kabir Peay.
The Committee was Kimberly G. Smith, Chair, University of Arkansas; Elisabeth 
Arevalo, Providence College; Erin Kuprewicz, National Museum of Natural 
History; and Kyle Harms, Louisiana State University.  The Committee would like 
to thank all the students that submitted packets for consideration.  "This year 
we received the most nominations and the best nominations" said Smith.  
"Choosing the winner this year was a difficult task given the quality of the 
nominations."


Kimberly G. Smith
Distinguished Professor of Biological Sciences
Department of Biological Sciences
University of Arkansas
Fayetteville, AR 72701
Phone:  479-575-6359  fax: 479-575-4010
Email:  [email protected]


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