For students, faculty, personnel from federal and state agencies, museums, 
environmental organizations and consulting firms

Taxonomy and Biology of Ferns and Lycophytes
August 14 - 20

Instructors: Robbin Moran and Carl Taylor
Location: Eagle Hill Institute, Steuben, Me

This course covers the identification, phylogeny, and ecology of ferns and 
lycophytes. Morning lectures will review the major families and place these in 
a phylogenetic context. The afternoon field trips will emphasize identification 
and ecology of local genera and species. The course will visit several habitat 
types along and near the eastern Maine coast to see as many species as 
possible. Herbarium specimens of northeastern species not found locally will be 
available for study. Besides identification, we will discuss the distinctive 
biology of ferns, such as their life cycle, hybridization, polyploidy, unusual 
adaptations (iridescent ferns, ant ferns, apogamy), biogeography, and uses of 
ferns by people. We will also discuss the sweeping changes that have taken 
place over the past ten years in understanding the phylogeny of ferns and 
lycophytes (for instance, why the term “fern allies” is no longer valid). We 
will discuss the biology and identification of quillworts (Isoëtes), a group in 
which Carl is a world expert. Plan to get your feet wet and consider bringing 
challenging specimens.

Dr. Robbin Moran ([email protected]) is Curator of Ferns at The New York 
Botanical Garden. He has published four books and over 80 scientific papers on 
ferns. With Alice Tryon, he has written the Ferns and Allied Plants of New 
England (1997) and with Barbara Joe Hoshizaki, the Fern Grower’s Manual (2001). 
He published A Natural History of Ferns (2004), which covers many aspects of 
fern biology and will be used as a text for the course. Robbin has traveled 
widely in Latin America and taught fern courses in Costa Rica, Venezuela, 
Ecuador, and Bolivia. During summers he co-teaches Tropical Plant Systematics, 
a five-week long course for in Costa Rica sponsored by the Organization for 
Tropical Studies.

Dr. W. Carl Taylor ([email protected]) is a retired museum curator of 
botany. Currently, he is a Research Associate at the Smithsonian Institution in 
Washington, DC. As a boy, Carl became interested in ferns through Boughton 
Cobb’s Ferns and Fern Allies and its wonderful illustrations by Laura Louise 
Foster. He served as Secretary of the American Fern Society for many years and 
published many papers in the American Fern Journal. Most of his articles are 
about Quillworts (Isoëtes), a curious lycophyte that looks like a tuft of 
grass. Carl has adventured far and wide by car, boat, helicopter, and even 
horseback in search of his beloved Quillworts. He is the author of Arkansas 
Ferns and Fern Allies.

For general information, go to 
http://eaglehill.us/programs/nhs/natural-history-seminars.shtml

For course calendar and course descriptions, go to 
http://www.eaglehill.us/programs/nhs/nhs-calendar.shtml

For application information and cost breakdown, go to 
http://www.eaglehill.us/programs/general/application-info.shtml

For more information, contact [email protected], 207-546-2821 x 1 

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