Don,

I was thinking more along the line of periodic flooding being a controlling 
factor in which species survive in a niche rather than a vector for seed 
dispersal.  Dale and I were talking about a paper on the Allegheny River Island 
stuff we are doing.  There are several distinct zones as you move upward in 
elevation from the normal river level to higher grounds.  on the lowest levels 
you find species that tolerate frequent floods sycamores, silver maples, black 
willow.  A little higher you begin to get more black locust, hawthorn and 
basswood,  A little higher and you get red oaks added to the mix, some hickory 
and butternut.  This progression represents the frequency of flooding for those 
different elevations.  Finally at the highest elevations in areas that are 
flooded only in the couple hundred year range you can pick up pines and 
hemlocks and a much wider diversity of species.  The major and most noxious 
invasives are limited in number or absent from the areas that are flooded every 
year or at least every other year.   At higher elevations which flood less 
frequent;ly, thanks to the wondrous Kinzua Dam the invasive multiflora roses 
and knotweed are the only species that seem to be present in the understory of 
trees in many areas.  Certainly you are not getting any regeneration of the 
trees in these areas.

Ed


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  ----- Original Message ----- 
  From: DON BERTOLETTE 
  To: [email protected] 
  Sent: Thursday, December 11, 2008 3:20 PM
  Subject: [ENTS] Re: Shrub-layer an empty niche?


  Ed-
  Your comments about invasive grasses set me to thinking...some of what you 
were saying has to do with the relative periodicity of the disturbances...if 
their return cycles go outside of their natural range of variation, the site 
may become more susceptible to invasives...flood events are perfect vectors for 
many species and are indiscriminant to the native/invasive seeds carried.
  -DonRB
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