I believe this article addresses the questions you have.  Depending on the 
configuration of the connector Exchange will resolve or not resolve the sender 
address.

http://technet.microsoft.com/en-us/library/bb232021.aspx

Thanks,
   Peter Dahl.

From: Campbell, Rob [mailto:[email protected]]
Sent: Tuesday, October 20, 2009 12:31 PM
To: MS-Exchange Admin Issues
Subject: RE: SMTP oddities with Exchange 2007

I'm not sure if I'm following all that exactly, but it sounds like you should 
be able to create a new receive connector that you can test (via telnet, PS 
script, etc) from one of your existing application servers by adding an 
additional IP address to the Exchange server, and binding the new receive 
connector to just that IP address.




From: Doug Gallimore [mailto:[email protected]]
Sent: Tuesday, October 20, 2009 11:21 AM
To: MS-Exchange Admin Issues
Subject: SMTP oddities with Exchange 2007

I have this weird issue with email sent from application boxes through the SMTP 
relay in a receive connector I setup with 2k7.  The sending address is an 
alias/proxy address on a public folder.  When the mail is received, the "from" 
address is "resolved" if you will to the public folder name, and not the proxy 
address, unless you view the header.  Apparently, a bunch of folks used to have 
rules that processed the messages based on the from address, and another 
application automatically creates tickets based on the from address, and it 
can't examine the header.  If I do a manual SMTP message through the hub 
server, I get the correct from, but the to, is blanked.  If I include the 
To:/From: in the DATA portion of the SMTP message it's all good, however, if I 
use the same test from telnet on one of the application servers, the from 
address gets "resolved" again.  I think it's a setting in my receive connector 
that is doing this, but I've no idea which one, my connector is pretty open to 
lots of internal application servers (mostly monitoring applications) which I 
have little control over.  So I was just going to create a new receive 
connector and lock it down to just one auth and one permissions group until I 
find it, but I'm having trouble isolating a test application server that isn't 
"critical".   Any suggestions?

Douglas

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