Once you get a handle to the file, via the File Class, you can use 
nativePath property to find an operating system specific path to the file:

  http://livedocs.adobe.com/apollo/1.0/aslr/flash/filesystem/File.html

  However, based on previous posts in this thread I was under the 
impression this would not work for you.

zdenekmikan wrote:
> 
> 
> Yes, it is the internal application, and I am just looking for that
> better solution - how to get UNC paths from File > browseForOpen method.
> 
> --- In [email protected] <mailto:flexcoders%40yahoogroups.com>, 
> Tom Chiverton <[EMAIL PROTECTED]>
> wrote:
>  >
>  > On Monday 14 Apr 2008, zdenekmikan wrote:
>  > > on PC. To be able to get access to these files from Mac AIR app I need
>  > > the name of server which is mapped to this O: letter. And to be able
>  > > to access files uploaded from Mac on PC I need to know the letter on
>  > > which the server with saved name is mapped on PC.
>  >
>  > I'm not sure I want an AIR app reading those sort of details of my
> internal
>  > network.
>  > If this is an internal app (sounds like it) there is probably a better
>  > solution (like common mount points, or using UNC paths directly with
> no drive
>  > letter).
>  >
>  > --
>  > Tom Chiverton
>  > Helping to appropriately streamline wireless technologies
>  > on: http://thefalken.livejournal.com <http://thefalken.livejournal.com>
>  >
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-- 
Jeffry Houser
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AIM: Reboog711  | Phone: 1-203-379-0773
--
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