In Nature today

http://www.nature.com/news/v-is-for-vortex-1.14507,
http://www.nature.com/nature/journal/v505/n7483/full/505295a.html, and
http://www.nature.com/nature/journal/v505/n7483/full/nature12939.html

describe instrumenting juvenile northern bald ibis raised in captivity to
measure their effort as they learn to fly in a V formation.  When they get
it right, the wing strokes are staggered in time so each bird, except the
leader, gets a lift off the trailing vortex of the bird ahead.

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