Compiling and linking these two files together should succeed.

/* BEGIN 1.cpp */
int callme();
int main(int argc, char** argv) { return callme(); }
/* END 1.cpp */

/* BEGIN 2.cpp */
template<typename T> struct holder { static int var; };
template<typename T> int holder<T>::var = 0;
int callme() { return holder<void>::var; }
/* END 2.cpp */

Indeed, compiling and linking them together without LTO succeeds. However, with
LTO, linking fails because of an undefined reference:

$ g++-4.5 -flto 1.cpp 2.cpp
/tmp/ccORg22M.lto.o: In function `main':
ccsrvz5g.o:(.text+0x10): undefined reference to `_Z6callmev'
collect2: ld returned 1 exit status

           Summary: Static members of templates cause LTO to omit required
           Product: gcc
           Version: 4.5.1
            Status: UNCONFIRMED
          Severity: normal
          Priority: P3
         Component: lto
        AssignedTo: unassigned at gcc dot gnu dot org
        ReportedBy: alexey at feldgendler dot ru
 GCC build triplet: x86_64-linux-gnu
  GCC host triplet: x86_64-linux-gnu
GCC target triplet: x86_64-linux-gnu

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