Hello,
I've noticed an inexplicably large difference between the
D.melanogaster 15-way phastcons data available for download in the
fixed step format
(in particular,
http://hgdownload.cse.ucsc.edu/goldenPath/dm3/phastCons15way/chr2L.pp.gz)
and the same in the variableStep format accessible from the Tables
view for the same D.melanogaster assembly.

They do look very similar at the beginning of the chromosome, as
expected, - for example here are the first 20:

pos_on_chr2L   variableStep    fixedStep
1       0.770953        0.771
2       0.77874 0.78
3       0.770953        0.772
4       0.77874 0.785
5       0.77874 0.78
6       0.755378        0.761
7       0.716441        0.722
8       0.708654        0.71
9       0.708654        0.716
10      0.700866        0.703
11      0.708654        0.712
12      0.700866        0.701
13      0.661929        0.667
14      0.59963 0.604
15      0.475031        0.479
16      0.264772        0.27
17      0.140173        0.146
18      0.132386        0.138
19      0.101236        0.109
20      0.101236        0.103

But by as early as 5400 they already look completely out of sync:

pos_on_chr2L   variableStep    fixedStep
5400    0.958661        0.251
5401    0.966331        0.345
5402    0.966331        0.383
5403    0.966331        0.39
5404    0.950992        0.376
5405    0.935654        0.229
5406    0.904976        0.211
5407    0.851291        0.166
5408    0.743921        0.152
5409    0.690236        0.116
5410    0.582866        0.044
5411    0.575197        0.017
5412    0.552189        0.019
5413    0.490835        0.017
5414    0.368126        0.01
5415    0.368126        0.018
5416    0.345118        0.022
5417    0.291433        0.039
5418    0.184063        0.05
5419    0.130378        0.054
5420    0.10737 0.054

This goes well beyond the listed loss of resolution for this fragment
(Worst case: 0.00760937).

>From some functional tests it seems that the variableStep data makes
more sense - although I've got not solid proof.
Can you please look into this - maybe I'm just confusing something?

Many thanks
Mikhail

--
Mikhail Spivakov, PhD
Postdoctoral Fellow
EMBL-EBI
Hinxton
Cambridge CB10 1SD
UK
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