The file I use to get all the colors the way I like them is "gtk.css". I don't know where one puts such a file for a Linux installation, but I've attached the file. The section you're looking for is grouped under "Register Settings" or something like that.

I'm running Gnucash 4.14, by the way. Hopefully my settings will be pertinent for you.
/*
 * Author: Kenneth J. Farley
 * Date:   2021-12-11
 *
 * GTK3 CSS file for use with GnuCash.
 * Developed by looking at complete theme definitions,
 * and scouring the internet for documentation.
 */

/* --- [ Variable Definitions ] ------------------------
 *
 * Definitions of colors so they can be used in many
 * places and subsequently changed conveniently, rather than
 * trying to find all uses of a particular color, etc.
 */

@define-color hueClear          rgba(   0,   0,   0, 0.00 ) ;
@define-color hueBGButton       rgba( 240, 240, 240, 1.00 ) ;
@define-color hueBGDropdown     rgba( 250, 235, 215, 1.00 ) ;
@define-color hueBGMain         rgba( 220, 220, 220, 1.00 ) ;
@define-color hueBGNormal       rgba( 220, 220, 220, 1.00 ) ;
@define-color hueBGScroll       rgba( 250, 250, 250, 0.40 ) ;
@define-color hueBGSlider       rgba(  72,  61, 139, 1.00 ) ;
@define-color hueBGTooltip      rgba(   0,   0,   0, 0.80 ) ;
@define-color hueBGTrough       rgba( 150, 150, 150, 0.80 ) ;
@define-color hueBGSelected     rgba(   0, 128, 128, 1.00 ) ;
/*
 * Hover   Custom Orange  = 255, 136,  77
 *         Chartreuse     = 127, 255,   0
 *         Carolina Blue  =  87, 160, 211
 *         Neon Green     =  57, 255,  20
 */
@define-color hueHover          rgba( 255, 136,  77, 1.00 ) ;
@define-color hueHoverLo        rgba( 255, 136,  77, 0.10 ) ;
@define-color hueHoverMd        rgba( 255, 136,  77, 0.20 ) ;
@define-color hueHoverHi        rgba( 255, 136,  77, 0.50 ) ;
@define-color hueHoverBorder    rgba( 255, 136,  77, 1.00 ) ;
/*
 * Tabs
 */
@define-color hueTabHover       rgba(  26, 140, 255, 1.00 ) ;
@define-color hueTabHoverLo     rgba(  26, 140, 255, 0.10 ) ;
@define-color hueTabHoverMd     rgba(  26, 140, 255, 0.20 ) ;
@define-color hueTabHoverHi     rgba(  26, 140, 255, 0.50 ) ;
@define-color hueTabHoverBorder rgba(  26, 140, 255, 1.00 ) ;
/*
 * Shade   LightCyan      = 224, 255, 255
 *         LightSteelBlue = 176, 196, 222
 *         LightYellow    = 255, 255, 214
 */
@define-color hueShadedLo       rgba( 176, 196, 222, 0.05 ) ;
@define-color hueShadedMd       rgba( 176, 196, 222, 0.20 ) ;
@define-color hueShadedHi       rgba( 176, 196, 222, 0.50 ) ;
/*
 * Hilite  White          = 255, 255, 255
 */
@define-color hueHiliteLo       rgba( 255, 255, 255, 0.10 ) ;
@define-color hueHiliteMd       rgba( 255, 255, 255, 0.25 ) ;
@define-color hueHiliteHi       rgba( 255, 255, 255, 0.50 ) ;
@define-color hueBorderNormal   lightsteelblue ;
@define-color hueProgress       rgba(   0, 102, 102, 1.00 ) ;
@define-color hueRegCursor      rgba( 230, 195,   0, 1.00 ) ;
@define-color hueRegDark        lightsteelblue ;
@define-color hueRegHeader      darkslateblue ;
@define-color hueRegLight       lemonchiffon ;
@define-color hueTextNormal     black ;
@define-color hueTextInverted   white ;

/*
 * --- [ Overall Appearance ] --------------------------
 *
 * Settings that will affect all occurrences of the
 * indicated components, unless they are overriden later.
 */

*
{
  font-family:      "Lucida Grande", sans-serif ;
  font-size:        11pt ;
}

button
{
  background-color: @hueBGButton ;
  background-image: linear-gradient(@hueShadedLo,@hueShadedMd,@hueShadedHi) ;
  border-radius:    6px ;
  border-style:     solid ;
  border-color:     @hueBGNormal ;
  border-width:     1px ;
}

button:hover
{
  background-color: @hueBGButton ;
  background-image: linear-gradient(@hueHoverLo,@hueHoverMd,@hueHoverHi) ;
  border-style:     solid ;
  border-color:     @hueHoverBorder ;
  border-width:     1px ;
}

/*
 * --- [ Status Bar ] ----------------------------------
 *
 * The bar at the bottom of the frame window that gives
 * a brief bit of info like the file name, and also holds
 * the progress bar that shows status of a read or save.
 */

statusbar label
{
  font-size:        11pt ;
  font-weight:      bold ;
  color:            @hueTextNormal ;
}

progressbar
{
  background-color: @hueClear ;
  font-size:        10pt ;
  margin:           0px 2px 2px 0px ;
}

progressbar.horizontal trough,
levelbar.horizontal trough,
progressbar.horizontal progress
{
  min-height:        10px ;
  border-radius:     5px ;
}

progressbar progress
{
  background-color: @hueProgress ;
  background-image: linear-gradient(@hueHiliteLo,@hueHiliteMd,@hueHiliteHi) ;
  height:           auto ;
  border-style:     none ;
}

/*
 * --- [ Main Window Bottom Bar ] ----------------------
 *
 * The small bar at the bottom of the application that
 * is visible for all operations.
 */

.GncMainWindow
{
  background-color: @hueBGMain ;
}

/*
 * --- [ Summary Bar ] ---------------------------------
 *
 * The "button" at the bottom of the Accounts page that
 * shows the total worth, profit, etc.
 */

.GncAccountPage
{
  background-color: @hueBGMain ;
}

.summary-bar
{
  background-color: @hueClear ;
}

.summary-bar cellview
{
  background-color: @hueClear ;
  color:            @hueTextNormal ;
}

.combo
{
  background-color: @hueBGButton ;
  background-image: linear-gradient(@hueShadedLo,@hueShadedMd,@hueShadedHi) ;
  color:            @hueTextNormal ;
  font-size:        16pt ;
  font-weight:      bold ;
}

/*
 * --- [ Notebook ] ------------------------------------
 *
 * The Notebook is the GUI structure Gnucash uses to show
 * the list of accounts, account registers, etc. It is a
 * simulation of a tabbed set of files. The settings here
 * will format those tabs, giving them similar behavior
 * as that for buttons (highlighting mouse hover, etc.)
 */

.GncMainWindow notebook box > image
{
  margin-left:      -2em ;
  margin-right:     -2em ;
  padding:          0px ;
  opacity:          0 ;
}

tabs, tab
{
  margin:           2px ;
  padding:          2px ;
}

tabs label
{
  margin:           2px ;
}

notebook > header
{
  background-color: @hueBGMain ;
  border-color:     @hueBGMain ;
}

notebook tab
{
  background-color: @hueBGButton ;
  background-image: linear-gradient(@hueShadedLo,@hueShadedMd,@hueShadedHi) ;
  min-height:       32px ;
  margin:           1px 1px 0px 0px;
  padding:          0px ;
  border-style:     solid ;
  border-color:     @hueBGNormal ;
  border-width:     1px 1px 0px 1px ;
  border-radius:    6px 6px 0px 0px ;
}

notebook tab:hover
{
  background-image: linear-gradient(@hueTabHoverLo,@hueTabHoverMd,@hueTabHoverHi) ;
  border-color:     @hueTabHoverBorder ;
}

notebook tab:checked
{
  border-color:     @hueTabHoverBorder ;
}

notebook > header.left tabs tab
{
  margin:           0px ;
  padding:          0px ;
  border-style:     solid ;
  border-color:     @hueBGNormal ;
  border-width:     1px 0px 1px 1px ;
  border-radius:    6px 0px 0px 6px ;
}

notebook > header.left tabs tab:hover
{
  border-color:     @hueTabHover ;
}

notebook >header.left tabs tab:checked
{
  border-color:     @hueTabHoverBorder ;
}

notebook tab box
{
  padding:          0px ;
  margin:           0px ;
  background-color: @hueClear ;
  border-width:     0px ;
}

notebook > header tab button:hover,
notebook > header tab button:hover:checked
{
  background-image: linear-gradient(@hueTabHoverLo,@hueTabHoverMd,@hueTabHoverHi) ;
  border-color:     @hueTabHoverBorder ;
}

/*
 * --- [ Scroll Bars ] ---------------------------------
 *
 * The scroll bars that show up on data presentations that
 * are too big for the current window.
 */

scrollbar
{
  background-image: none ;
  border-style:     solid ;
  background-color: @hueBGScroll ;
}

scrollbar trough
{
  border-style:     none ;
  border-width:     0px ;
  background-color: @hueBGTrough ;
}

scrollbar.vertical trough
{
  background-image: linear-gradient(to right,@hueHiliteLo,@hueHiliteMd,@hueHiliteHi) ;
}

scrollbar slider
{
  background-color: @hueBGSlider ;
}

/*
 * --- [ Account Tree ] --------------------------------
 *
 * The list of accounts, usually the first window tab
 * in the program.
 */

#account_tree
{
  letter-spacing:   0px ;
  background-color: lavender ;
  color:            @hueTextNormal ;
}

.GncAccountPage box
{
  background-color: @hueClear ;
}

.GncAccountPage label
{
  font-weight:      bold ;
  background-color: @hueClear ;
  color:            @hueTextInverted ;
}

#account_tree:selected
{
  background-color: @hueBGSelected ;
  color:            @hueTextInverted ;
  font-weight:      bold ;
}

treeview button
{
  background-color: @hueBGNormal ;
  color:            @hueTextNormal ;
}

#account_tree header button
{
  background-color: @hueRegHeader ;
  background-image: linear-gradient(@hueShadedLo,@hueShadedMd,@hueShadedHi) ;
  color:            @hueTextInverted ;
  border-radius:    0px ;
  border-style:     solid ;
  border-color:     @hueBGNormal ;
  border-width:     0px 1px 0px 0px ;
}

#account_tree header button:hover
{
  background-image: linear-gradient(@hueHiliteLo,@hueHiliteMd,@hueHiliteHi) ;
}

/*
 * --- [ Toolbar Settings ] ----------------------------
 *
 * These settings change the appearance of the toolbar
 * and its buttons.
 */

toolbar
{
  background-color: @hueBGMain ;
}

toolbar toolbutton button
{
  background-color: @hueBGButton ;
  background-image: linear-gradient(@hueShadedLo,@hueShadedMd,@hueShadedHi) ;
  padding-left:     2px ;
  padding-right:    2px ;
  border-color:     @hueBorderNormal ;
  min-width:        64px ;
}

toolbar toolbutton button:hover
{
  background-color: @hueBGButton ;
  background-image: linear-gradient(@hueHoverLo,@hueHoverMd,@hueHoverHi) ;
  border-color:     @hueHoverBorder ;
}

toolbar toolbutton label
{
  font-size:        11pt ;
  padding-left:     2px ;
  padding-right:    2px ;
}

/*
 * --- [ Register Settings ] ---------------------------
 *
 * Set the alternating colors for the lines of an account
 * register, the cursor (line being edited), the split
 * lines, etc.
 */

.GncRegisterPage
{
  background-color: @hueBGMain ;
}

.GncRegisterPage label
{
  font-family:      "CopperPlate", sans-serif ;
  font-size:        12pt ;
  background-color: @hueClear ;
  color:            @hueTextNormal ;
  margin:           2px ;
}

*.gnc-class-register-header
{
  background-color: @hueRegHeader ;
  color:            @hueTextInverted ;
}

*.gnc-class-register-primary
{
  background-color: @hueRegLight ;
  color:            @hueTextNormal ;
}

*.gnc-class-register-primary:disabled
{
  background-color: mix (@hueRegLight, grey, 0.2) ;
  color:            mix (@hueTextNormal, grey, 0.2) ;
}

*.gnc-class-register-secondary
{
  background-color: @hueRegDark ;
  color:            @hueTextNormal ;
}

*.gnc-class-register-secondary:disabled
{
  background-color: mix (@hueRegDark, grey, 0.2) ;
  color:            mix (@hueTextNormal, grey, 0.2) ;
}

*.gnc-class-register-split
{
  background-color: @hueBGDropdown ;
  color:            @hueTextNormal ;
}

*.gnc-class-register-split:disabled
{
  background-color: mix (@hueBGDropdown, grey, 0.2) ;
  color:            mix (@hueTextNormal, grey, 0.2) ;
}

*.gnc-class-register-cursor
{
  background-color: @hueRegCursor ;
  color:            @hueTextNormal ;
}

*.gnc-class-register-cursor:disabled
{
  background-color: mix (@hueRegCursor, grey, 0.2) ;
  color:            mix (@hueTextNormal, grey, 0.2) ;
}

gnc-id-cursor entry
{
  margin:           2px 5px 2px 5px ;
  background-color: @hueRegCursor ;
  color:            @hueTextNormal ;
}

gnc-id-cursor button
{
  margin:           1px 1px 1px 1px ;
}

/*
 * --- [ Tooltips ] ------------------------------------
 *
 * These are the small popups that give a brief line of text
 * explaining the function of a button, etc.
 */

tooltip.background
{
  background-color: @hueClear ;
}

tooltip box
{
  background-color: @hueBGTooltip ;
  color:            @hueTextInverted ;
  border-radius:    4px ;
  padding:          0px 0px 0px 0px ;
}

/*
 * --- [ Calendar ] ------------------------------------
 *
 * Apparently someone thought the calendar was important,
 * since they provided a lot of means of altering its
 * appearance.
 */

sheet calendar
{
  font-family:      "Arial", sans-serif ;
  font-size:        10pt ;
}

calendar
{
  border-color:     @hueHoverBorder
}

calendar.header
{
  background-color: @hueBGNormal ;
  border-radius:    2px ;
  border-width:     1px ;
  border-color:     @hueBGNormal ;
  background-image: linear-gradient(@hueShadedLo,@hueShadedMd,@hueShadedHi) ;
  color:            @hueTextNormal ;
}

calendar.highlight
{
  background-color: @hueBGSelected ;
  color:            @hueTextInverted ;
  border-radius:    0px ;
  padding:          0px ;
}

calendar.view
{
  background-color: @hueRegDark ;
  background-image: linear-gradient(to right,@hueHiliteLo,@hueHiliteHi) ;
  color:            @hueTextNormal ;
}

#dens-cal-popup treeview
{
  background-color: @hueBGDropdown ;
}

calendar.button
{
  background-color: @hueBGButton ;
  background-image: linear-gradient(@hueShadedLo,@hueShadedMd,@hueShadedHi) ;
  border-radius:    2px ;
  border-style:     solid ;
  border-color:     @hueBGNormal ;
  border-width:     1px ;
}

calendar.button:hover
{
  background-color: @hueBGButton ;
  background-image: linear-gradient(@hueHoverLo,@hueHoverMd,@hueHoverHi) ;
  border-style:     solid ;
  border-width:     1px ;
  border-color:     @hueHoverBorder ;
}

calendar*:selected,
calendar*:selected:focus
{
  background-color: @hueBGSelected ;
  color:            @hueTextInverted ;
  border-radius:    2px ;
}
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