Personally I think its related to a code splitting bug that has been fixed 
in GWT 2.6. The bug causes the GWT compiler / code splitter to not emit 
some needed defineSeed() statements in split points. The symptoms mentioned 
in the below issue/discussion are not 100% your symptoms but I think it is 
somewhat related.

https://code.google.com/p/google-web-toolkit/issues/detail?id=7392
https://groups.google.com/forum/#!msg/google-web-toolkit-contributors/wCBk0qrM4n8/Rzami92kQfgJ

If you use GWT 2.5 and you are curious then temporary remove split points 
in your app. I would guess your issue will disappear.


-- J.

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