I then look up the MX record for the domain and find that Imail is trying to send to the "A" record and not the MX record!
Do you have 2 or more DNS servers listed in the IMail SMTP settings? If so, you should make sure that they are all working properly. Otherwise, if there is a temporary problem reaching the first one, IMail will go to the second one. But if the second one reports no MX record (because the IMail server isn't authorized to do lookups), and IMail then checks the A record at the first server (which it *should* do if the MX record does not exist), IMail will go to the A record.
-Scott
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