We would like to draw attention to our new paper, which develops 
and uses new methods (mixture models) for analyzing mark-
recapture population data in which there is heterogeneity. The 
abstract is below and a pdf is available at:

http://whitelab.biology.dal.ca/hw/Whitehead_Wimmer_2005.pdf

Hal Whitehead and Tonya Wimmer 

Whitehead, H., and T. Wimmer. 2005. Heterogeneity and the 
mark-recapture assessment of the Scotian Shelf population of 
northern bottlenose whales (Hyperoodon ampullatus). Canadian 
Journal of Fisheries and Aquatic Sciences 63:2573-2585.

A population of northern bottlenose whales (Hyperoodon 
ampullatus) inhabits the waters along the edge of the Scotian 
Shelf. The most important habitat of this population is the Gully, a 
large submarine canyon, where animals were photographically 
identified between 1988 and 2003. Open mark–recapture models, 
including mixture models that allow for heterogeneity in 
identifiability and (or) mortality among individuals, were fitted to 
identification-history data. Models without heterogeneity in 
identifiability had poor fit to the data and underestimated population 
size. The population is estimated to contain about 163 animals 
(95% confidence interval 119–214), with no statistically significant 
temporal trend. About 12% of the population has a high probability 
of being identified within the Gully in any year. Many of them are 
mature males. The remainder is less likely to be identified in the 
Gully during any year, spend generally shorter periods in the Gully 
even in years when they are found, and are more likely to be 
female. This and other work indicate a poorly mixed population 
inhabiting the canyons and other deeper waters off the Scotian 
Shelf. Nonparametric bootstrap methods were used to validate the 
estimation procedure and to estimate the efficiency of future  
fieldwork scenarios.

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