Dear colleagues,

heres' a new paper that might be of interest. Please also note my new working 
address....(out of Germany, finally.....). For those having questions or for 
assistance, please refer to me directly.

Best,

Frank


Structural and temporal emission patterns of variable pulsed calls in 
free-ranging killer whales (Orcinus orca) 
Authors: Rehn, Nicola; Teichert, Stefanie; Thomsen, Frank

Source: Behaviour, Volume 144, Number 3, 2007, pp. 307-329(23)

Abstract:

Resident killer whales off Vancouver Island, British Columbia, produce variable 
burst pulsedcalls most commonly during close-range interactions such as 
socialising or social-travelling. Earlier studies indicated that variable calls 
are graded and can be arranged into a scale from low-frequency calls to 
high-frequency ones. These graded calls are often emitted in sequences, were 
call-classes of similar frequency follow one another more often than different 
classes. However, a detailed analysis of sequences was lacking to date. 
Therefore, our understanding of the function of variable calls during 
interactions among killer whales is rather limited. Simultaneous recordings of 
underwater vocalizations and behavioural observations from resident killer 
whales were collected off Vancouver Island, British Columbia during1996-2001. 
Socialising activities were divided into four categories: male-female, 
male-male, female-juvenile and juvenile-juvenile. Variable call sequences were 
analy!
 sed with RTS and SIGNAL acoustic-software. We found no positive correlation 
between group-size and number of used calls or the duration of sequences, 
indicating that only one or a few animals were involved in the production of 
each sequence. Furthermore, sequences were present in all four behaviour 
categories and the composition of the group had no influence on the duration of 
calls and used call-classes. One particular call class (V4) could be further 
separated into structurally distinct sub-classes. These sub-classes often 
formed rather stereotyped sequences. The results of our study indicate that 
sequences of variable calls emit broad motivational information that is not age 
or sex-related. Sequences of distinct sub-classes might encode more subtle 
information on emotional states during socialising. Therefore, variable calls 
might posses different functions, depending on the nature of the interaction. 
Thus, variable calls might be of great importance for close-range comm!
 unication in wild killer whales. 

Dr. Frank Thomsen
Cefas
Regulatory Assessment Team
Burnham Laboratory
Remembrance Avenue
Burnham-on-Crouch
Essex CMO 8HA
Tel :  ++ 44 (0) 1621 787 227
Fax : ++ 44 (0) 1621 784 989
Mobile: ++ 44 (0) 7984586499
www.cefas.co.uk




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