The following paper is now available online at Inter-Research Marine Ecology 
Progress Series; full text is available at 
http://www.int-res.com/abstracts/meps/v373/p173-186/, or by emailing me 
([email protected]):
 
Resolving fine-scale spatio-temporal dynamics in the harbour porpoise Phocoena 
phocoena 
Henrik Skov and Frank Thomsen
 
Abstract: 
With the development of human activities in offshore shelf waters, the harbour 
porpoise Phocoena phocoena has become a focal species in relation to management 
of human activities and marine habitat conservation efforts. In this study, 
acoustic and visual time series of harbour porpoise occurrence were analysed 
from the Horns Reef area, eastern North Sea, aiming at resolving periodicity, 
spatial scale of aggregation and habitat drivers in porpoises. A total of 51 
fine-scale surveys and 474 d of hydrophone deployment at 2 stations were 
analysed by partial least squares regression and spatial modelling in relation 
to synoptic (spatio-temporal) data derived from a local, high-resolution 
hydrodynamic model. Small-scale changes in local currents reflecting upwelling 
driven by the interaction of the semi-diurnal tidal currents with the steep 
slopes of the bank were the main habitat driver identified from the acoustic as 
well as the survey data. Spatial modelling based on selected key predictor 
variables showed the distribution of harbour porpoises to alternate between 2 
upwelling cells depending on the direction of tidal currents. The size of 
upwelling cells was less than 10 km, matching the small-scale spatial structure 
of survey data as revealed by geostatistical analysis. The findings may have 
great implications for the design of future protected areas for harbour 
porpoises, and they indicate that the overlap between areas of concentration of 
the species and areas targeted by fisheries and the offshore wind industry 
could be avoided without major changes to current management practices. 



Henrik Skov
DHI Water & Environment
Agern Alle 5
DK-2970 Hørsholm
Denmark

Telephone (45)45169220
Cell/Private (46)702478477
Fax (45)45169292
Skype henrikskov7249
www.dhigroup.com
 


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