Dear Marmam subscribers,
 
The following are abstracts from one of the most recent issues (Volume 13, 
issue 2, 2013) of the Journal of Cetacean Research and Management. The 
following is posted on behalf of the IWC and the journal editor.
 
The International Whaling Commission (IWC) publishes the Journal of Cetacean 
Research and Management thrice yearly (Spring, Autumn, and Winter), with at 
least one supplement that will contain the full report of the IWC Scientific 
Committee. 
This is one of the final issues of JCRM to be published in hard copy format. 
The IWC is delighted to inform you that the Journal of Cetacean Research and 
Management will be available free of charge online from Volume 14 
(https://iwc.int/jcrm). Back copies will also be available free of charge on 
the IWC website. Some back copies of the Journal of Cetacean Research and 
Management are available in hard copy format, and these are available for the 
cost of postage only. For further details, please contact [email protected].
 
Contact information is provided for the corresponding author for each article. 
Please do not contact the listserve editors or me for pdfs or copies of the 
articles. Thank you for your continued interest in the journal and abstract 
postings. 
A guide for authors is included at the website.
 
With regards,
 

Dagmar Fertl
Sumich, J.L., S.A. Blokhin, and P.A. Tiupeleyev. 2013. Revised estimates of 
foetal and post-natal growth in young gray whales (Eschrichtius robustus). 
Journal of Cetacean Research and Management 13(2):89-96.
 
Contact e-mail: [email protected]
 
Patterns of gray whale growth in body length and weight with age are evaluated 
using published values of foetal and post-natal body dimensions at reported 
ages, supplemented with previously unpublished measured lengths of 88 
mid-gestation foetuses, 82 first-summer calves and 30 secondsummer whales taken 
in the summer/autumn Chukotkan native subsistence fishery. Gompertz growth 
models are fitted to foetal and post-natal lengths at age, predicting mean 
lengths at birth in mid-January of 4.7m, 7.9m at weaning and 8.7m at one year. 
The late foetal diapause in growth of length is not supported by the available 
data. Two equations were derived for estimating body weights from the linear 
body dimensions of length nd maximum girth. For biomass estimates, two 
equations based on length alone and on both girth and length are derived. A 
multiple least squares regression equation fit to 14 measurements of the same 
whale over 14 months of captive rehabilitation predicts mean body weights at 
birth of 1,100–1,200kg, 5,100–5,200kg at six months (weaning), and 
6,700–6,800kg at one year of age.
******
D’Intino, A.M., J.D. Darling, J. Urbán R., and T.R. Frasier. 2013. Lack of 
nuclear differentiation suggests reproductive connectivity between the 
‘southern feeding group’ and the larger population of eastern North Pacific 
gray whales, despite previous detection of mitochondrial differences. Journal 
of Cetacean Research and Management 13(2):97-104.
 
Contact e-mail: [email protected]
 
During winter, eastern North Pacific gray whales migrate south to calving 
grounds in the lagoons of Baja California, and in spring they migrate north to 
their summer feeding grounds in the Chukchi and Beaufort Seas. Although the 
majority of the population makes this migration, a small subset of the 
population known as the ‘southern feeding group’ ends their northward migration 
early, spending summers feeding in waters ranging from northern California to 
southern Alaska. Previous analyses based on photo-ID and mtDNA data indicate 
that this seasonal substructuring results from maternally-directed site 
fidelity to different feeding grounds, and that this site fidelity and feeding 
ground preference is passed from mothers to their offspring. It is currently 
assumed, but not known, that the individuals of the southern feeding group mate 
with the rest of the population, and therefore that the eastern North Pacific 
gray whale represents one interbreeding population. Testing this assumption and 
understanding how these whales are related to the rest of the population, is 
key to making appropriate management decisions, which are particularly relevant 
given the recent increase in potential removals, or threats in the area such as 
the proposed resumption of aboriginal whaling, and increased oil pipeline 
development and subsequent vessel traffic. This paper analyses 15 nuclear 
microsatellite loci in 59 individuals from the southern feeding group and 40 
individuals from the calving lagoons (representative of the larger population) 
to test the hypothesis that the eastern North Pacific gray whale represents one 
interbreeding population. No indication of population substructuring was found 
based on these nuclear loci, suggesting that all sampled whales do indeed 
represent one interbreeding population. Combined, these data from mitochondrial 
and nuclear markers therefore suggest one interbreeding population that is 
seasonally subdivided based on maternally-directed site fidelity to different 
feeding areas.
****
Coughran, D.K., N.J. Gales, and H.C. Smith. 2013. A note on the spike in 
recorded mortality of humpback whales (Megaptera novaeangliae) in Western 
Australia. Journal of Cetacean Research and Management 13(2):105-108.
 
Contact e-mail: [email protected]
 
An unprecedented number of humpback whales (n = 46) from Breeding Stock D (BSD) 
were found dead or dying on Western Australian beaches in 2009. This compares 
to an average of less than four stranded humpback whales in each year in the 
period between 1989 and 2008. The recorded number of humpback whales on beaches 
in the year preceding this peak (2008, n = 13) and the two years following the 
peak (2010, n = 16; 2011, n = 17) were also above the long term average. In 
2012 (n = 7), the numbers of stranded whales was closer to the longer term 
average. The majority of stranded individuals were either calves (44%) or 
juveniles/sub-adults (49%), with only 7% classified as adults. Most whales 
appeared emaciated. There were insufficient data to determine cause of death. 
Three possible hypotheses to explain this spike in mortality are proposed: (1) 
the increase in mortality of BSD was an artefact of increased detection and 
reporting; (2) the increase was temporary and transitory; and (3) the spike in 
mortality represents the start of an increasing trend in mortality as the 
population approaches carrying capacity. We suggest that hypothesis two is the 
most plausible but on-going monitoring will be required to test this.
*****
 
Rankin, R.W., D. Maldini, and G. Kaufman. 2013. Bayesian estimate of Australian 
humpback calving interval under sparse resighting rates: 1987-2009. Journal of 
Cetacean Research and Management 13(1):109-121.
 
Contact e-mail: [email protected]
 
This study estimates a calving interval for humpback whales from a longterm 
photo-ID catalogue of 2,973 individuals resighted in Hervey Bay, East 
Australia. The study proposes a modification of two existing methods to handle 
partial identification of sex and age-classes of whales from visual surveys. 
One method truncates the data to just breeding females and discards all 
resighting events prior to the first observed breeding event. The second method 
utilises the multi-stage mark recapture (MSMR) framework and multi-event 
extension to include all resighted individuals and their entire encounter 
history. The performance of each method is assessed and the conditioning 
required to handle ambiguity of sex and age-classes is detailed, which is 
subtly different from most other mark-recapture methods. Both truncation and 
the multi-event methods led to similar estimates of calving intervals: 2.98 
years (95% CI: 2.27–3.51) and 2.78 years (95% CI: 2.23–3.68) respectively. More 
importantly, estimates were more sensitive to the exact specification of 
resighting probabilities among age and sex classes than to the type of 
conditioning. However, the multi-event framework resulted in more precise 
estimates of other important life-history parameters such as apparent survival, 
and included a wider constituency of age and sex classes.
*****
Hakamada, T., K. Matsuoka, S. Nishiwaki, and T. Kitakado. 2013. Abundance 
estimates and trends for Antarctic minke whales (Balaenoptera bonaerensis) in 
Antarctic Areas IV and V for the period 1989/90-2004/05. Journal of Cetacean 
Research and Management 13(2):123-151.
 
Contact e-mail:[email protected]
 
The Japanese Whale Research Programme under Special Permit in the Antarctic 
(JARPA) conducted sighting surveys during the 1989/90 to 2004/05 austral summer 
seasons (mainly in January and February), alternating between IWC management 
Areas IV (70°E–130°E) and V (130°E–170°W), both south of 60°S each 
(split-)year. These data are analysed to obtain abundance estimates for 
Antarctic minke whales (Balaenoptera bonaerensis) in these Areas. The estimates 
are calculated by standard line transect analysis methods using the program 
DISTANCE under the assumption that g(0) = 1. Annual rates of increase in 
abundance are estimated using log-linear models. The analyses take several 
recommendations from the 2006 JARPA Review Meeting into consideration. Those 
addressed here aim to: (a) improve the point estimates of abundance and their 
precision; and (b) evaluate (through sensitivity tests) the effect of different 
factors associated with the JARPA survey on the estimates of abundance and 
trend. GLM models are used to adjust for different strata being surveyed at 
different times of year over the duration of JARPA, with model selection being 
based on AICc. Abundance estimates for Area IV range from 16,562 (CV = 0.542) 
in 1997/98 to 44,945 (CV = 0.338) in 1999/00, while those for Area V range from 
74,144 (CV = 0.329) in 2004/05 to 151,828 (CV = 0.322) in 2002/03. Estimates of 
the annual rates of increase in abundance are 1.8% with a 95% CI of [–2.5%, 
6.0%] for Area IV and 1.9% with a 95% CI of [–3.0%, 6.9%] for Area V. Estimates 
of these trends are robust to the effects of changes in survey timing, the 
shapes of the shoulders of detection functions, portions of survey tracklines 
following the ice edge, parts of the Areas in which no survey took place and 
poor coverage within some strata. Adjustments to allow for the g(0) being less 
than 1 are made by the application of a regression model, developed from the 
results of the Okamura-Kitakado (OK) method estimate of minke whale abundance 
from the IDCR-SOWER surveys, which provides estimates of g(0) from the 
statistics of the minke whale school size distribution in a stratum. With this 
adjustment, abundance estimates increase by an average of 32,333 (106%) for 
Area IV and 89,245 (86%) for Area V, while the estimates of annual rates of 
increase and their 95% CIs change slightly to 2.6% [–1.5%,6.9%] for Area IV and 
1.6% [ 3.4%,6.7%] for Area V.
*****
Murase, H., I. Temoai, T. Kirata, S. Finkaso, G. Yasunaga, and L.A. Pastene. 
2013. A note on cetaceans off Kiribati and Tuvalu from a research cruise in 
October 2010. Journal of Cetacean Research and Management 13(2):153-158.
 
Contact e-mail: [email protected]
 
This paper summarises the results of a sighting survey conducted around the 
Gilbert Islands (Kiribati) and Tuvalu Islands (Tuvalu) in the central Pacific 
Ocean between 3 and 17 October 2010. This was the first systematic collection 
of cetacean sighting data in this region. The main objective of the survey was 
to investigate the occurrence and distribution of cetaceans around Kiribati and 
Tuvalu. In addition biopsy samples were obtained to investigate the species 
identity of Bryde’s-whale-like baleen whales through genetic analyses and to 
assess feeding ecology of cetaceans in the survey area through the examination 
of fatty acids. The survey was carried out using a sighting survey vessel, 
which covered a total of 1,012 n.miles (≈1,875km). A total of 24 schools (640 
individuals) of cetaceans was sighted: three schools (five individuals) of 
Bryde’s-whale-like baleen whales, one school (nine individuals) of sperm whales 
(Physeter macrocephalus), one school (six individuals) of killer whales 
(Orcinus orca), one school (14 individuals) of short finned pilot whales 
(Globicephala macrorhynchus), one school (two individuals) of false killer 
whales (Pseudorca crassidens), eight schools (483 individuals) of spinner 
dolphins (Stenella longirostris), and one school (70 individuals) of striped 
dolphins (Stenella coeruleoalba). Mitochondrial DNA (mtDNA) analyses based on 
the biopsy samples identified two of the Bryde’s-whale-like whales sighted, as 
of the putative species Balaenoptera brydei. Compositions of fatty acids of 
Bryde’s and killer whales are presented. The survey provided new information on 
the distribution and fatty-acid composition of cetaceans around Kiribati and 
Tuvalu.
*****
Stevick, P.T., J.M. Allen, M.H. Engel, F. Félix, B. Haase, and M.C. Neves. 
2013. Inter-oceanic movement of an adult female humpback whale between Pacific 
and Atlantic breeding grounds off South America. Journal of Cetacean Research 
and Management 13(2):159-162.
 
Contact e-mail: [email protected]
 
We report the first documented movement of an individual humpback whale between 
the eastern South Pacific stock off Ecuador and the western South Atlantic 
stock off Brazil. This constitutes the first record of a humpback whale in both 
the Atlantic and Pacific breeding grounds off South America, and one of a small 
number of inter-oceanic movements documented to date. It is possible that, even 
at quite low levels, this movement of individuals between breeding grounds 
contributes to the current high level of mtDNA diversity in these once-depleted 
Southern Hemisphere populations. When first sighted, the whale was accompanied 
by a young calf, and is therefore identified as an adult female. This movement 
to a different and distant breeding ground is the first reported by a 
reproductively mature female, and shows that extreme long-distance travellers 
among humpback whales are not restricted to young males.
*****
Iriarte, V., and M. Marmontel. 2013. Insights on the use of dolphins (boto, 
Inia geoffrensis and tucuxi, Sotalia fluviatilis) for bait in the piracatinga 
(Calophysus macropterus) fishery in the western Brazilian Amazon. Journal of 
Cetacean Research and Management 13(2):163-173.
 
Contact e-mail: [email protected]
 
In the Amazon Basin, the use of the pink dolphin or boto (Inia geoffrensis) for 
bait in the piracatinga (Calophysus macropterus) fishery was first detected in 
the year 2000. Since then, this artisanal fishery has become more prevalent as 
it requires only a few hours of work per night and provides immediate cash 
earnings. It is thus an attractive addition to (or replacement for) traditional 
fishing. Previous reports have noted the use of botos as bait, but stated that 
the most common bait used are caimans (Melanosuchus niger, Caiman crocodilus). 
Estimates of the number of dolphins killed based on fish landings have been 
proposed and an apparent decrease in sighting/survival of an 
artificially-marked boto population was observed. Although stocks/population 
estimates, trends and actual numbers of hunted dolphins are unknown, the 
conservation impacts of this activity are of concern. Between October 2010 and 
November 2011, research was conducted within an area with serious conflicts 
between dolphins and fishermen as well as intense fishing for piracatinga, i.e. 
in the lower Japurá River, on the border with the Mamirauá and Amanã 
Sustainable Development Reserves, where both boto and tucuxi (Sotalia 
fluviatilis) are used for bait. One-hundred and fifty-seven monitoring surveys 
were carried out in eight key communities, confirming 114 piracatinga fishing 
events through direct monitoring and incognito surveys of fishing gear 
(gaiolas). Empirical evidence of the activity in gaiolas comprised pieces of 
bait, carcass remains, piracatinga provoked vomits and dolphin fished 
carcasses. Of those, 31.2% (n = 35) involved cetacean bait (91.4% I. 
geoffrensis, 8.58% S. fluviatilis), 68.7% (n = 77) caiman bait (96% M. niger, 
4% C. crocodilus), and two fishing events used both types. These percentages 
may be higher/lower in other areas within and outside the Reserves. Given the 
increasing trend of the piracatinga fishery, the authors believe that 
precautionary measures for the conservation of Amazonian dolphins are urgently 
needed. Development of practical short-term solutions (e.g. offal-baited fish 
traps) and multispecies management together with law enforcement, incentives 
and educational programmes could allow the future transition of riverine 
communities from the piracatinga fishery to sustainable, higher income 
activities.
*****
Ryan, C., D. Craig, P. López-Suárez, J.V. Perez, I. O’Connor, and S.D. Berrow. 
2013. Breeding habitat of poorly studied humpback whales (Megaptera 
novaeangliae) in Boa Vista, Cape Verde. Journal of Cetacean Research and 
Management 13(2):175-180.
 
Contact e-mail address: [email protected]
 
The waters surrounding Cape Verde comprise one of two known breeding grounds 
for humpback whales in the North Atlantic. The population remains very small 
and has apparently failed to recover since the cessation of whaling there. 
During the breeding seasons of 2011 and 2012, sighting surveys were carried out 
for humpback whales off Boa Vista, the easternmost island of the Cape Verde 
Island archipelago. The distribution and relative abundance of humpback whales 
and mother-calf pairs was investigated by plotting effort-corrected sightings 
using a 2km2 grid-square. The study area, a 206km2 region from the coastline up 
to 8km offshore, covered the western half of Boa Vista where whales have 
previously been regularly recorded. Following 1,954km of search effort, 117 
sightings of humpback whales were made. An encounter rate of 0.11 whales per km 
was recorded for both years. It is hoped that these data may assist in 
implementing conservation measures to protect humpback whales and the habitat 
of Baia Sal Rei, which appears to be the single most important bay for winter 
breeding, calving and nursing humpback whales in the eastern North Atlantic.





























































 
                                          
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