Two potential PhD Studentship Opportunities: (1) Grey seal behaviour, (2) 
Application of biotelemetry to quantify behaviour.

Below are details of two potential PhD opportunities being advertised at Durham 
University (UK). Both are competitive studentships, and all initial enquiries 
should be made to Dr  Sean Twiss 
([email protected]<mailto:[email protected]>)



PROJECT 1: Behavioural and physiological responses to changing environments in 
wild grey seals (Halichoerus grypus)

Supervisory team:

Dr. Sean Twiss (Durham Univ.): 
[email protected]<mailto:[email protected]>

Dr. Patrick Pomeroy (SMRU, University of St. Andrews)

Dr Amanda Bishop (Alaska SeaLife Center)



Background: Organisms have evolved mechanisms to cope with natural stressors, 
but rapid environmental change is subjecting species to new threats or more 
unpredictable and extreme natural stressors. Conventional studies focus on 
species or ecosystem level responses to change, such as range or phenological 
shifts, but these are products of variation in individual responses. Within 
populations, individuals can show differing coping-styles; proactive 
individuals express little flexibility and are less responsive to environmental 
stimuli, whereas reactive individuals are more flexible and responsive. 
Understanding how individuals differ in their ability to cope with stressors, 
and the consequences if they fail to do so, is critical for quantifying 
species’ resilience to current and future threats of climate change and 
anthropogenic disturbance.



This project aims to investigate variation in behavioural and physiological 
stress reactivity within and between individuals, and across populations, using 
extensive archived data on known individual adult female grey seals 
(Halichoerus grypus) during the breeding season, a time of intense energetic 
demands and limitations.  Our previous studies have highlighted the 
considerable additional stress placed on breeding grey seals by extreme weather 
patterns, and this study will examine changing behavioural patterns in concert 
with indicators of physiological changes in breeding and non-breeding wild grey 
seals. The study will assess whether behavioural indicators of environmental 
stress map onto physiological indicators, and establish levels of 
inter-individual variation, and within individual consistency  in such 
responses.


Methods: The study will utilise an extensive existing catalogue of data on 
known individual breeding grey seals from a range of contrasting breeding 
colonies. The successful candidate will extract new data from archived video 
footage of these same seals to provide key behavioural and physiological 
metrics. These metrics will be analysed with respect to prevailing 
environmental factors, including social context (e.g. conspecific density) and 
weather conditions. Data analysis will involve mixed effects modelling 
procedures using ‘R’. There may also be scope for some additional field-based 
data collection, but this will depend upon opportunity.

Candidate requirements: Potential candidates should be able to demonstrate the 
ability to conduct prolonged periods of behavioural observations, while 
remaining focused, dedicated and enthusiastic. Ability to work independently 
and as part of a team is essential. In addition, candidates will require strong 
analytical skills, including experience of modern ecological and statistical 
modelling techniques. Experience with R and analysis of behavioural data is 
advantageous, but not essential.  There is the possibility of additional data 
collection through field campaign(s), but candidates will need to show a 
willingness and ability to travel and conduct prolong field observations alone. 
A UK driving licence (or equivalent) would be essential for field campaigns.

Training: The student will gain extensive inter-disciplinary training in 
behavioural observation, video analysis, and data analysis in R employing 
mixed-effects models. The student will also develop critical thinking, writing, 
presentational and teamwork skills as part of dynamic, interdisciplinary, and 
supportive research groups.



Eligibility

Applications for this project are welcome from suitably qualified candidates 
worldwide. Applicants must be applying to start a full-time PhD in the Faculty 
in October 2020. They must have fulfilled the University’s requirements 
regarding English language ability and must not require a presessional course 
as a condition of their place.

See: https://www.dur.ac.uk/learningandteaching.handbook/1/3/3/



Funding and how to apply

This project is in competition with others for funding (via the Durham Doctoral 
Studentship scheme). Success will therefore depend on the quality of 
applications received, relative to those for competing projects. If you are 
interested in applying, in the first instance contact Dr Sean Twiss, 
([email protected]) with a CV and covering letter, detailing your reasons 
for applying for the project. Only the best applicants will be asked to submit 
a full application. UK and Non-UK students areadvised that to be considered for 
the Durham Doctoral Fellowship, the minimum requirements include an excellent 
undergraduate degree with either postgraduate or work experience in a relevant 
discipline and scientific publication(s) in peer reviewed journals.



Application deadline: Friday 10th Jan. 2020 at 5 pm (GMT)



-------------------------------------------------------------------------------------


PROJECT 2: Quantifying the relative importance of extrinsic and intrinsic 
drivers of individual behaviour and habitat use of hefted sheep on a biodiverse 
but fragile upland ecosystem.

Although this project does not involve seals, it does require a range of 
research skills that marine mammalogists often possess; experience of the 
deployment of biotelemetry devices and analysis of biotelemetry data to examine 
behaviour patterns in free ranging mammals.

Supervisory team:

Dr. Sean D. Twiss ([email protected]<mailto:[email protected]>), 
Department of Biosciences, Durham University

Dr. Robert Baxter, Department of Biosciences, Durham University

Martin Furness, Senior Reserve Manager, Natural England

Dr Mike Morecroft, Natural England and University of Oxford

Dr Gavin Stewart, Newcastle University

Dr Fiona Lovatt, School of Veterinary Science, University of Nottingham.


Background:   The unenclosed upland areas of the UK are internationally 
important for wildlife and of high nature conservation value. However, these 
fragile habitats are subject to various threats including the impacts of 
climate change, atmospheric pollution deposition, acid rain and grazing 
pressures. A primary tool for sustainable management of these landscapes is 
implementing grazing regimes that allow maintenance or recovery of habitats and 
which support ecosystem services.  However, remarkably little is known about 
the behaviour of the major grazer of UK uplands, domestic hill sheep, and their 
impact on vegetation mosaics. This project aims to investigate the determinants 
of group and individual level foraging choices within upland vegetation mosaics 
and how these vary in relation to sheep age, size, experience (hefting) and 
extrinsic factors including the spatial distribution of vegetation types, 
weather and micro-topography. This study will provide valuable information on 
how these different factors interact and allow a consideration of practical 
management techniques that can be used to influence the spatial distribution 
and feeding choices of sheep.

Upper Teesdale National Nature Reserve hosts unique Arctic-Alpine plant 
assemblages with many species existing in climatically marginal locations. This 
project will involve extensive fieldwork at Widdybank Fell, part of the NNR 
that contains some of the most biodiverse habitat, and for which pre-existing 
fine-scale habitat maps exist. Sheep behaviour will be monitored at the level 
of the individual using a combination of in situ visual observation, 
supplemented by video recordings, and by using animal borne telemetry; GPS 
data-loggers will be deployed to provide fine scale movement and location data, 
with accelerometers to allow for automatic classification of behaviours with 
respect to time and location. The research will require the development of 
analytical protocols to classify raw accelerometry data into behavioural 
categories, including foraging.  Behavioural data will be used within a GIS to 
examine intrinsic and extrinsic drivers of individual sheep behaviour and 
habitat use. Widdybank Fell is a long established research site and benefits 
from detailed existing data and ongoing studies on micrometeorology, 
microtopography and microhabitat. The current study will have the potential to 
integrate behavioural data with these environmental data within a GIS.

The research will involve extensive field based behavioural observations of 
individually identified sheep. Potential candidates should be able to 
demonstrate the ability to conduct prolonged fieldwork in potentially harsh 
conditions, while remaining dedicated and enthusiastic. Ability to work 
independently and as part of a team is essential. In addition, candidates will 
require strong analytical skills, including experience of modern ecological and 
statistical modelling techniques. Experience with R and analysis of telemetry 
data is advantageous, but not essential.  A UK driving licence (or equivalent) 
is essential. Further details of the project and candidate requirements can be 
obtained from Dr. Twiss.

The student will gain extensive interdisciplinary training in fieldwork 
logistics, behavioural observation, deployment of telemetry devices, and 
develop analytical skills for both behavioural and telemetry data. The student 
will also develop critical thinking, communication, and teamwork skills as part 
of dynamic and interdisciplinary research groups.


This project is in competition with others for funding, and success will depend 
on the quality and suitability of applicants, relative to those for competing 
projects. For further information, or to apply, contact Dr. Sean Twiss at 
[email protected]. In your email include: 1) two-page covering letter 
explaining your reasons for applying and why you selected this project, 2) CV 
with contact information for two references, 3) Full transcripts of previous 
qualifications. Only the best applicants will be asked to submit an application 
to the University. The application deadline is 10th January 2020, therefore 
students should contact Dr. Twiss well in advance for initial consideration. 
For eligibility requirements, see http://www.iapetus.ac.uk/aboutstudentships/



END


_________________________________

Dr. Sean Twiss,
Associate Professor in Animal Behaviour and Behavioural Ecology,
Department of Biosciences,
South Road,
Durham University,
Durham, DH1 3LE,
UK.

E-mail: [email protected]
Web-site: 
https://www.dur.ac.uk/biosciences/about/schoolstaff/academicstaff/?id=1132
Blog: http://sealbehaviour.wordpress.com/
Behaviour, Ecology and Evolution Research (BEER) Centre: 
www.dur.ac.uk/beer-centre<https://owa.dur.ac.uk/owa/redir.aspx?C=AjvknJfcq0-zlL0498uhGGKmvrw4G9MIOkl7uzB2o0DQWJkijfaedd4PLox8gN2oJ64a8h9XCa8.&URL=http%3a%2f%2fwww.dur.ac.uk%2fbeer-centre>

Tel: +44 (0)191 334 1350 (office)
Tel: +44 (0)191 334 1247 (lab)
Fax: +44 (0)191 334 1201
_________________________________
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