Dear Marmam and ECS members (sorry for crossposting),

on behalf of my coauthors, I am pleased to inform you on our recent publication 
in Nature Scientific Reports. We show that deep-diving pilot whales can 
echolocate almost continuously whilst foraging at depth by using tiny amounts 
of air to make each click and by capturing and recycling the used air. The 
small air volumes mean that echolocation doesn’t take much energy.

The full reference and abstract are found below:

Foskolos, I., Aguilar de Soto, N., Madsen, P.T. & Johnson, M. Deep-diving pilot 
whales make cheap, but powerful, echolocation clicks with 50 µL of air. Sci. 
Rep. 9, 15720 (2019).

Abstract
Echolocating toothed whales produce powerful clicks pneumatically to detect 
prey in the deep sea where this long-range sensory channel makes them 
formidable top predators. However, air supplies for sound production compress 
with depth following Boyle’s law suggesting that deep-diving whales must use 
very small air volumes per echolocation click to facilitate continuous sensory 
flow in foraging dives. Here we test this hypothesis by analysing click-induced 
acoustic resonances in the nasal air sacs, recorded by biologging tags. Using 
27000 clicks from 102 dives of 23 tagged pilot whales (Globicephala 
macrorhynchus), we show that click production requires only 50 µL of air/click 
at 500 m depth increasing gradually to 100 µL at 1000 m. With such small air 
volumes, the metabolic cost of sound production is on the order of 40 J per 
dive which is a negligible fraction of the field metabolic rate. Nonetheless, 
whales must make frequent pauses in echolocation to recycle air between nasal 
sacs. Thus, frugal use of air and periodic recycling of very limited air 
volumes enable pilot whales, and likely other toothed whales, to echolocate 
cheaply and almost continuously throughout foraging dives, providing them with 
a strong sensory advantage in diverse aquatic habitats.

This is an Open Access paper and it is freely accessible online at: 
https://www.nature.com/articles/s41598-019-51619-6.

If you have any problems with the above link or any questions, please feel free 
to contact me at [email protected]<mailto:[email protected]>.

Best wishes,
Ilias

Ilias Foskolos, PhD Fellow
Marine Bioacoustics Lab<https://marinebioacoustics.wordpress.com/>

Zoophysiology, Dept. Bioscience
Aarhus University
C.F. Møllers Allé 3, Building 1131
DK-8000 Aarhus C, Denmark
E-mail: [email protected]<mailto:[email protected]>

ResearchGate<https://www.researchgate.net/profile/Ilias_Foskolos3>
Twitter: theclickitclick<https://twitter.com/theclickitclick>
Phone: (+45) 50656572

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