Dear MarMamers
For those interested in the heart rate response during diving, this article may 
be of interest and summarizes the current knowledge of the capacity of 
vertebrates to cognitively alter heart rate. The abstract and doi are included 
below and the link provides a number of free downloads. If you would like a pdf 
copy and cannot access it below, please send me an email.
Sincerely,
Andreas

Title: The role of cognition as a factor regulating the diving responses of 
animals, including humans

Authors: Fahlman, A., Burggren, W., Milsom, W.K.

Abstract: The dive response involves three main components - breath holding, 
reduced heart rate and increased peripheral vasoconstriction - and is 
ubiquitous during forced dives in air-breathing vertebrates; however, numerous 
studies in free-diving animals have shown that the heart rate response to 
diving varies considerably in a manner that suggests cognitive control. 
Furthermore, studies on free-diving animals and controlled experiments in 
trained animals both indicate that the dive response can be conditioned, such 
that the reduction in heart rate begins before submergence and the extent of 
the reduction is set early in the dive. In addition, numerous species also 
experience an increase in heart rate and blood flow during ascent at the end of 
a dive, a phenomenon commonly called 'ascent tachycardia'. Collectively, these 
data suggest that although the dive response is under autonomic control, many 
species can vary its magnitude depending on the length and type of the planned 
dive - an indication of a role for cognition in the overall physiological 
responses associated with diving. Here, we provide examples of the conditioned 
cardiac responses - including anticipatory changes in heart rate - in several 
diving species and propose potential underlying mechanisms. We also discuss how 
the anticipatory cardiovascular responses not only improve diving capacity, but 
also prevent diving-related problems, such as decompression sickness or 
barotrauma, through a mechanism described by the selective gas exchange 
hypothesis.

doi: https://doi.org/10.1242/jeb.246472

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