My co-authors and I are pleased to announce our recent publication in The ISME 
Journal titled Respiratory microbiomes reflect whale health:

Carolyn A Miller and Enrico Pirotta, Sharon Grim, Michael J Moore, John W 
Durban, Peter L Tyack, Holly Fearnbach, Samantha G M Leander, Amy R Knowlton, 
Amy M Warren, Monica A Zani, Regina Asmutis-Silvia, Heather M Pettis, Amy 
Apprill, Respiratory microbiomes reflect whale health, The ISME Journal, Volume 
19, Issue 1, January 2025, wraf231, https://doi.org/10.1093/ismejo/wraf231

Abstract
As important members of the marine ecosystem, baleen whales are frequently 
managed and protected, but methodology to assess their health remains limited. 
Recent technological advances, such as the use of drones, support the 
non-invasive collection of promising health-associated data, including 
respiratory exhalant microbiota. Here, we considered five health metrics paired 
with respiratory exhalant samples to examine the utility of characterizing 
respiratory microorganisms for health diagnostics of North Atlantic right 
whales (Eubalaena glacialis), one of the most endangered baleen whale species. 
In 2016–2024, we used drones to collect 103 exhalant samples from 85 
individuals to examine the associated microbiome, using amplicon sequencing 
methods targeting bacteria and archaea. The health status of sampled whales was 
characterized using an index of body condition derived from full-body vertical 
drone images, three qualitative assessments obtained from photo-identification 
imagery, and an existing health and vital rates model. Using an elastic net 
penalized regression approach, we demonstrate significant relationships between 
these health metrics and respiratory-associated microorganisms. Bacterial taxa 
that significantly contributed to the model for the body condition index 
differed between the thinnest and most robust males in the dataset. The thin 
whale harbored taxa belonging to the same genus as mammalian pathogens, 
Clostridium and Peptoniphilus, whereas the robust whale harbored taxa commonly 
observed in lipid-rich environments, Sediminispirochaeta and Candidatus 
Gracilibacteria. These differences warrant further investigation into the 
mechanisms by which bacteria contribute to whale health. Our findings 
demonstrate the utility of non-invasive multi-metric health models that include 
respiratory exhalant microbiota for whale health assessment and management.

The paper is available through open access at:
https://academic.oup.com/ismej/article/19/1/wraf231/8319970

Kind regards,
Carolyn Miller

Carolyn A. Miller, Ph.D.
Woods Hole Oceanographic Institution
Department of Marine Chemistry and Geochemistry
Mail Stop #4
266 Woods Hole Road
Woods Hole, MA 02543
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