New publication on rough-toothed dolphin social network structure:

Corsi, E., R.W. Baird, S.D. Mahaffy, J.J. Kiszka. 2026. Social life in the 
deep: variation in network structure and association patterns of rough-toothed 
dolphins (Steno bredanensis) around the main Hawaiian Islands. Marine Mammal 
Science 42:e70281.  https://doi.org/10.1111/mms.70281

Abstract
While coastal delphinid sociality has been extensively investigated across the 
globe, little is known about the social structure of odontocetes in the open 
ocean. The rough-toothed dolphin (Steno bredanensis) is a globally distributed 
oceanic dolphin occurring in tropical and warm-temperate waters. Around the 
main Hawaiian Islands, two demographically distinct populations have been 
identified: one off Kauaʻi and Niʻihau, the other off Hawaiʻi Island. We 
investigated differences in network structure and association patterns between 
these populations using photo-identification data collected from 2002-2020. 
Social network analysis revealed that the Kauaʻi–Niʻihau population was more 
subdivided into distinct social clusters (i.e., had higher modularity: 0.58 vs. 
0.47) and less densely interconnected (i.e., lower network density: 0.07 vs. 
0.12) than the Hawaiʻi Island population. Mean association strength within 
clusters was higher off Hawaiʻi Island (0.31) than off Kauaʻi–Niʻihau (0.22). 
Social connections off Hawaiʻi Island also relied more strongly on a smaller 
number of individuals that linked clusters together. We suggest that 
differences in population density and habitat extent may contribute to 
differences in social network structure. Although low re-sighting rates limited 
our ability to characterize either population comprehensively, applying 
identical methods allowed us to identify relative differences in network 
structure between them.

If you would like a pdf contact Enrico Corsi 
([email protected]<mailto:[email protected]>) or myself.

For more on what Cascadia Research does check out our end-of-summer wrap-up 
flipbook https://cascadiaresearch.aflip.in/Summer_2026.html

--
Robin W. Baird, Ph.D.
Hawai‘i Program Director
Cascadia Research Collective<https://cascadiaresearch.org/>
218 ½ W. 4th Avenue
Olympia, WA 98501 USA
https://orcid.org/0000-0002-9419-6336
Donate to support our work<https://cascadiaresearch.org/donations/>
Follow us on Facebook<https://www.facebook.com/CascadiaResearch/>
Updates from our August Kaua‘i field 
project<https://cascadiaresearch.org/hawaii-update/august2026/>

Recent publications<https://cascadiaresearch.org/hawaii_publications/>:

Baird, R.W. 2026. Dwarf and pygmy sperm whales Kogia sima (Owen, 1866) and 
Kogia breviceps (de Blainville, 1838). Pages 313-352 in Ridgway and Harrison's 
Handbook of Marine Mammals, Volume 3 River Dolphins and Larger Toothed Whales. 
Edited by T.A. Jefferson. Academic Press Elsevier.

Corsi, E., R.W. Baird, S.D. Mahaffy, and J.J. Kiszka. 2026. Social life in the 
deep: variation in network structure and association patterns of rough-toothed 
dolphins (Steno bredanensis) around the main Hawaiian Islands. Marine Mammal 
Science 42(4):e70281. https://doi.org/10.1111/mms.70281

Harnish, A.E., R.W. Baird, and M.A. Kratofil. 2026. Diving behavior of common 
bottlenose dolphins around the main Hawaiian Islands: individual variation and 
responses to diel and lunar cycles. Marine Mammal Science 42:e70232. 
https://doi.org/10.1111/mms.70232

Kratofil, M.A., J.F. Shaff, H.K. Hoffbauer, M. Cantor, M.C. Hill, and R.W. 
Baird. 2026. Ecological contexts of diving behavior in Hawaiian false killer 
whales. Movement Ecology https://doi.org/10.1186/s40462-026-00630-4

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