Well, the error message says you should check the SECRET_KEY setting.
Have you done that? Most likely it was removed from your settings module
when you upgraded to Mezzanine 4. I recommend you put it in
local_settings.py to avoid having it under version control.
Other than that, there's a know issue relating field injection of
non-Django fields and Django 1.9:
https://github.com/stephenmcd/mezzanine/issues/1513
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