I am glad that the correspondence between Igor and Dennis were made public 
and that the discussion about this topic continues. I think things have 
become much clearer for me.

I agree with Asta in almost all respects. Taxonomists like practical, 
diagnostic characters very much. It would be nice to see which characters 
(landmarks) are of taxonomic importance. But this is not always (and in 
fact, very often) the case. Let me put an example. Suppose there is an 
insect taxon having two distantly related food plants (this is very common 
situation). Or suppose there is a taxon with two discretely different wing 
patterns but otherwise being exactly similar (also very common situation). 
Suppose also that those forms are sympatric (live together in space and 
time). Now, the question is not what the differences are since they can 
easily been kept apart due to the discrete polymorphism. This polymorphism, 
however, does not imply they are valid species, since polymorphism is such 
features may be intraspecific. Thus the question is rather that does this 
polymorphism correlate with morphology (in insect this mean often genital 
morphology). The finding that the forms differ in genital morphology, 
regardless what the differences are, is of taxonomic relevance. It speaks 
for the co-existence of two biologically isolated species. In reference to 
Dennis's argument ("I believe the original question that started all of 
this was about discriminant functions. This is a bit different. If the goal 
of a discriminant function analysis is the classification of new specimens, 
then it, in a sense, matters not what you do. If the goal is 
classification, then the validity of whatever is done is determined solely 
by its ability to correctly classify new specimens. You could use the first 
few PCAs, every third partial warp, neural networks, or tarot cards. That 
which does the best job is the best of what you have tried") I therefore 
conclude that in my case selecting a qualified set of partial warps in 
order to classify cross-validated specimens is justified. As I found that 
cross-validated specimens were classified correctly, I conclude that there 
are differences, though perhaps indefinable, in analyzed genitalic outline 
sections, because such a result could not be achieved by chance. All this 
reasoning is basically founded on the assumption that difference in a 
single partial warp tells that there is(are) difference(s) in shape, even 
though this difference cannot be associated with any particular landmark. 
Correct? I also assume that by comparing the consensus configuration of 
forms with the overall consensus using thin-plate-splines and vector plots 
I can speculate what those differences possibly are.

Well, these are difficult things I am not sure my reasoning above makes 
sense. Therefore, I will be grateful for further comments, also critical 
ones. I also like to warmly thank all of you who have taken part to this 
discussion.

Cheers,

Marko


At 20:47 02/02/2005, you wrote:
>maybe this has been discussed already a lot but I would still dare to ask:
>
>To me it seems that for taxonomists, the use of individual partial warps
>comes largely from the need to interpret the CVA analysis and find the most
>informative characters to differentiate the species. These variables later
>can be used in further taxonomic work.
>
>In the "traditional" morphometrics one can add/remove variables one by one
>and leave the most informative ones - a widely used practice, I believe.
>
>With geometric morphometrics, for the mathematical reasons discussed before,
>one will conduct CVA on relative warp scores
>The results will tell that the most important variable for differentiating
>the two species is RW5. But this is not a variable a taxonomist would like
>to use. OK, I can go back to RW analysis and explore shape variation related
>to RW5. Still it is speculative, compared with the  rigorous statistical
>procedure of selecting "traditional" variables by their within to among
>group variance (or vice versa?).
>One can also remove a certain part of landmarks manually and do the analysis
>again, but this is also subjective.
>
>Probably it does not make much sense to choose the most important landmarks,
>because the shape is analysed as a whole, in relation to other landmarks?
>There may not be one or two "the most informative" landmarks, which could be
>informative alone?
>
>Yet, I wonder whether there is there any accepted statistical procedure to
>select the most useful characters (landmarks) for species differentiation?
>
>Thanks,
>
>Asta Audzijonyte
>
>
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>For more information visit http://www.morphometrics.org
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