-------- Original Message --------
Subject: Re: PLS and sources of covariance
Date: Tue, 5 Apr 2011 14:14:02 -0400
From: Chris Klingenberg <[email protected]>
Reply-To: [email protected]
Organization: University of Manchester
To: [email protected]

Dear Eric

Concerning this and previous posts: I guess the main problem is that the
relation between ontogenetic stage and shape is not linear. If so,
linear regression won't remove all of it. It might be possible that a
suitable transformation might render it linear -- the problem is finding
it... For allometry, log-transforming size often does a good job. So you
might get away with using a transformation for the measure of
ontogenetic stage that you use.

That litter mates cluster together can either be due to such differences
or to other maternal effects (physiology, nutrition, behaviour etc.; I
think you said you are using mice that are genetically identical, so
it's not genetic...).
Regardless of the cause of it heterogeneity, it might definitely be part
of the covariance among parts. To keep it constant, you could do a
pooled within-litter PLS (but that reduces your degrees of freedom to
the total number of embryos minus the number of litters).

I hope this helps.

Best wishes,
Chris



On 4/5/2011 6:22 PM, morphmet wrote:


-------- Original Message --------
Subject: PLS and sources of covariance
Date: Tue, 5 Apr 2011 10:57:25 -0400
From: [email protected]
To: [email protected]

Hi all,

I have a question regarding PLS and the interpretation of data.

My samples are mouse embryos, the heads and faces of midgestational staged
specimens. Shape coordinates are regressed on stage (number of somites) to
remove ontogenetic variation, and yet again on centroid size to remove
allometric variation (a multiple multivariate regression is probably more
appropriate, and I will try this). The general goal of these sorts of
regressions is to remove components of shape variation not related to
differences in genotype, so different genotypes can be compared.

The residual data are "blocked" into two regions that are subject to PLS
for an analysis of shape covariance between the two blocks. The goal of
the test is to remove ontogenetic and allometric variation, perform a PLS
comparing two separate blocks of landmarks, and look for associations that
are independent of ontogentic and allometric causes of between-block
covariance.

My question is related to my recent posts. I suspect that despite
regressing on stage and size, a lingering ontogenetic component remains in
the shape data. For example, in PCA scatter plots of the regressed data,
litter mates tend to cluster together. My question is that, if ontogenetic
variation persists in data treated as above, and a PLS test detects
significant shape covariance, is it not reasonable to suspect that the
cause of the relationship is due to the stubborn presence of variation
owing to ontogenetic development? The two blocks will appear to covary in
shape because the sample contains specimens that vary in developmental
progress and at least some of that variation is still present in the data
despite attempts to regress it out. Therefore, the two blocks will show a
relationship in the PLS test because some embryos will have blocks that
are each somewhat developmentally advanced and some embryos will have
blocks that are each less developmentally advanced, and this will create a
pattern of shape covariance.

Thanks.

Eric




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Christian Peter Klingenberg
Faculty of Life Sciences
The University of Manchester
Michael Smith Building
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United Kingdom

Telephone: +44 161 275 3899
Fax: +44 161 275 5082
E-mail: [email protected]
Web: http://www.flywings.org.uk
Skype: chris_klingenberg
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