Yep--they were officially split in 2002 or so, so if you've been down to
Black-crested habitat, you've got a lifer!

According to the Birds of North America Black-crested Titmouse entry:

Recent evidence from DNA-DNA hybridization (Slikas et al.
1996<http://bna.birds.cornell.edu/bna/species/717/articles/species/717/biblio/bib072>)
raised to full generic status the subgenera *Baeolophus* (New World titmice)
and *Poecile* (chickadees and related Old World tits) of the genus *Parus*,
a recommendation adopted by the American Ornithologists’ Union
(1997<http://bna.birds.cornell.edu/bna/species/717/articles/species/717/biblio/bib002>).
On basis of allozyme (Gill et al.
1989<http://bna.birds.cornell.edu/bna/species/717/articles/species/717/biblio/bib032>)
and DNA (Sheldon et al.
1992<http://bna.birds.cornell.edu/bna/species/717/articles/species/717/biblio/bib070>,
Slikas et al. 
1996<http://bna.birds.cornell.edu/bna/species/717/articles/species/717/biblio/bib072>)
data, *Baeolophus* is distantly related to other parids. As currently
recognized (American Ornithologists’ Union
1998<http://bna.birds.cornell.edu/bna/species/717/articles/species/717/biblio/bib003>),
*Baeolophus* is comprised of five species: the closely related Tufted and
Black-crested Titmice, the closely related Oak (*B. inornatus*) and Juniper
(*B. ridgwayi*) Titmice, and the enigmatic Bridled Titmouse (*B. wollweberi*).
The Tufted/Black-crested Titmouse clade is sister the to Oak/Juniper
Titmouse clade (Gill et al.
1989<http://bna.birds.cornell.edu/bna/species/717/articles/species/717/biblio/bib032>,
Sheldon et al. 
1992<http://bna.birds.cornell.edu/bna/species/717/articles/species/717/biblio/bib070>,
Slikas et al. 
1996<http://bna.birds.cornell.edu/bna/species/717/articles/species/717/biblio/bib072>
).

The Tufted and Black-crested Titmice, which are sister species, were merged
into a single species on the basis of free interbreeding in a narrow, stable
contact zone in central Texas (Allen
1907<http://bna.birds.cornell.edu/bna/species/717/articles/species/717/biblio/bib001>,
Dixon 
1955<http://bna.birds.cornell.edu/bna/species/717/articles/species/717/biblio/bib021>,
1978<http://bna.birds.cornell.edu/bna/species/717/articles/species/717/biblio/bib023>,
1989<http://bna.birds.cornell.edu/bna/species/717/articles/species/717/biblio/bib024>,
1990<http://bna.birds.cornell.edu/bna/species/717/articles/species/717/biblio/bib025>).
However, the American Ornithologists’ Union (Banks et al.
2002<http://bna.birds.cornell.edu/bna/species/717/articles/species/717/biblio/bib007>)
reevaluated genetic divergence (Braun et al.
1984<http://bna.birds.cornell.edu/bna/species/717/articles/species/717/biblio/bib009>,
Avise and Zink 
1988<http://bna.birds.cornell.edu/bna/species/717/articles/species/717/biblio/bib005>,
Sheldon et al. 
1992<http://bna.birds.cornell.edu/bna/species/717/articles/species/717/biblio/bib070>)
and evidence that vocalizations differ (Dixon
1955<http://bna.birds.cornell.edu/bna/species/717/articles/species/717/biblio/bib021>,
Coldren 
1992<http://bna.birds.cornell.edu/bna/species/717/articles/species/717/biblio/bib015>)
and concluded both that the taxa should be accorded species status and that
assortative mating prevailed in areas of contact.

Minor divergence (0.4 to 0.6%) in mitochondrial DNA (Avise and Zink
1988<http://bna.birds.cornell.edu/bna/species/717/articles/species/717/biblio/bib005>,
Gill and Slikas
1992<http://bna.birds.cornell.edu/bna/species/717/articles/species/717/biblio/bib031>)
implies that the Tufted and Black-crested Titmice have been isolated for
~250,000 yr, a finding that supports Dixon’s
(1978<http://bna.birds.cornell.edu/bna/species/717/articles/species/717/biblio/bib023>)
contention that the taxa became separated during the late Pleistocene. By
contrast, large protein (allozyme) divergence (*D* = 0.06; Gill and Slikas
1992<http://bna.birds.cornell.edu/bna/species/717/articles/species/717/biblio/bib031>)
suggests that the taxa are older and exchanged mtDNA through recent
hybridization in secondary contact.





On Fri, Mar 13, 2009 at 3:32 PM, Stan & Molly Jo Miller-Johnson <
[email protected]> wrote:

> In trying to update my bird list, I've discovered that there is now a
> Black-crested Titmouse.  Is it the same bird that used to be simply a
> version/morph of the Tufted Titmouse?  I'd be pretty happy if that was the
> case!  It would put me at 3 short of 500 life birds... unless as I continue
> this "official" count I find more that were split or merged!  Never a dull
> moment.
>
> I've looked on the internet in a couple places, but mostly I'm just seeing
> that the Black-crested is listed... but I'm not finding an official split
> off the Tufted.
>
> Thanks for any help you can provide!
>
> Molly Jo Miller
> Inver Grove Hts, Dakota Co.
>
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Twin Beaks!  www.twinbeaks.blogspot.com
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