The projection on the map is whatever the projection of current baselayer is. It will then request all other layers are served in this projection. The server providing the data should be reprojecting it on the fly. I am guessing the WMTS was your base layer. In general, 4326 is fine for storing data on your server, but as a "projection", (Plate Carree), it has no redeeming cartographic properties whatsoever.

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