I came across a Ted talk by Dan E with the title "How language began". At the halfway mark (9 minutes) he mentions Peirce and relates his semiotic to the issues: https://www.youtube.com/watch?v=qFxg5vkaPgk
My only comment would be that there was probably some kind of vocalization a few million years earlier. Chimpanzees in the wild use gestures to communicate among themselves. When Australopithecus began to walk upright, they could carry stuff (food, weapons, and babies). Sounds would be a useful supplement when their hands were full. John
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