Hello
   
  I realized
 Anova repeated mesur
And I have result following:
   
  summary(a)
  
Error: indiv
             Df Sum Sq Mean Sq F value  Pr(>F)    
Alt           2 28.377  14.188 63.5236 < 2e-16 ***
Site          9 75.238   8.360 37.4279 < 2e-16 ***
Espèce        1  1.002   1.002  4.4867 0.03514 *  
Competition   1  0.228   0.228  1.0202 0.31344    
Residuals   252 56.286   0.223                    
---
Signif. codes:  0 '***' 0.001 '**' 0.01 '*' 0.05 '.' 0.1 ' ' 1 
  Error: indiv:time
                  Df Sum Sq Mean Sq  F value    Pr(>F)    
time               3 68.328  22.776 762.3477 < 2.2e-16 ***
Alt:time           6  1.513   0.252   8.4387 6.998e-09 ***
time:Site         27  6.872   0.255   8.5192 < 2.2e-16 ***
time:Espèce        3  7.403   2.468  82.5979 < 2.2e-16 ***
time:Competition   3  0.121   0.040   1.3536    0.2559    
Residuals        756 22.586   0.030                       
---
Signif. codes:  0 '***' 0.001 '**' 0.01 '*' 0.05 '.' 0.1 ' ' 1 
> 

  But I do not know ( Error)?
   
   

                
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